Nature - 2019.08.29

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the tissue immunofluorescence staining. P.C. and S.H. performed bioinformatics
analysis. I.H., J.K. and A.O. performed the proteomics and analysed the data.
E.G.-G. helped with the collection of Ly6G+ cells for proteomics. G.M. performed
the 3D-scaffold co-culture to analyse CD104+ cells. A.W. and L.C. performed the
electron microscopy experiments. E.H. and V.S. provided human samples. L.O.,
E.N., I.K., V.B. and J.-H.L. critically reviewed the manuscript. J.-H.L. supervised the
lung organoid experiments. I.M. designed and supervised the study, interpreted
the data and wrote the manuscript.


Competing interests The authors declare no competing interests.


Additional information
supplementary information is available for this paper at https://doi.org/
10.1038/s41586-019-1487-6.
Correspondence and requests for materials should be addressed to J.-H.L. or
I.M.
Peer review information Nature thanks Marie-Liesse Asselin-Labat, Thomas
Tüting and the other, anonymous, reviewer(s) for their contribution to the peer
review of this work.
Reprints and permissions information is available at http://www.nature.com/
reprints.
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