Article
Foundation, Aquatic Symbiosis no. GBMF9198 (https://doi.org/10.37807/GBMF9198, Y.Z.),
NIH/NIGMS GM106023 (Y.Z.), GM110151 (Y.Z.), NIH/NIAMS AR060042 (C.-M.F.) and AR071976
(C.-M.F.).
Author contributions C.-M.F. and Y.Z. conceived and supervised the project. M.H., C.-M.F. and
Y.Z. designed experiments. M.H. performed the experiments. M.H. and X.Z. analysed the data.
M.H., X.Z., C.-M.F. and X.Z. interpreted the data and wrote the manuscript.
Competing interests The authors declare no competing interests.
Additional information
Supplementary information is available for this paper at https://doi.org/10.1038/s41586-020-
2385-7.
Correspondence and requests for materials should be addressed to M.H., C.-M.F. or Y.Z.
Peer review information Nature thanks Mónica Medina Munoz and the other, anonymous,
reviewer(s) for their contribution to the peer review of this work. Peer reviewer reports are
available.
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