Semicircular Canals
The three semicircular canalsare fluid-filled mem-
branous ovals oriented in three different planes. At the
base of each is an enlarged portion called the ampulla
(see Fig. 9–8), which contains hair cells (the crista) that
are affected by movement. As the body moves for-
ward, for example, the hair cells are bent backward
at first and then straighten (see Fig. 9–10). The bend-
ing of the hair cells generates impulses carried by the
vestibular branch of the 8th cranial nerve to the
cerebellum, midbrain, and temporal lobes of the cere-
brum. These impulses are interpreted as starting or
stopping, and accelerating or decelerating, or chang-
ing direction, and this information is used to maintain
equilibrium while we are moving (see Box 9–6: Motion
Sickness).
In summary then, the utricle and saccule provide
information about the position of the body at rest,
while the semicircular canals provide information
about the body in motion. Of course, there is some
overlap, and the brain puts all the information together
to create a single sense of body position.
The Senses 215
Utricle
Saccule
Otoliths
Hair cells
Gravity
Head tilted
8th cranial
nerve
Head upright
Ampulla
Crista
Ampulla
Hair
cells
8th cranial
nerve
1.
At rest
2.
Starting
3.
Moving
4.
Stopping
5.
At rest
A
B
Figure 9–10. Physiology of
equilibrium. (A) Utricle and
saccule. (B) Semicircular canals.
See text for description.
QUESTION: In part A, what
causes the hair cells to bend? In
part B, what causes the hair
cells to sway?