Encyclopedia of Biology

(Ron) #1

located at regular intervals along the actin filament and
allows actin to interact with myosin.
See alsoCYTOSKELETON.


actinomorphic Symmetrical over more than one ver-
tical plane; e.g., flowers that can be separated into sym-
metrical halves along any plane.


action potential A localized rapid change in voltage
that occurs across the membrane of a muscle or nerve
cell when a nerve impulse is initiated. It is caused by a
physicochemical change in the membrane during the
interaction of the flow and exchange of sodium and
potassium ions.
See alsoDEPOLARIZATION.


active center The location in an ENZYMEwherethe
specific reaction takes place.


active site The active site of an enzyme is the area—
a depressed region comprising a few of the protein’s
amino acids—on a portion of an enzyme that binds to
its substrate. The enzyme’s specificity is based on the
shape of the active area, which can alter itself to snugly
fit the substrate to which it is binding by using weak
chemical bonds.
SeeACTIVE CENTER.


active transport The movement of a substance
across a biological membrane, such as living cells,
against a concentration (diffusion) gradient with the
help of metabolic energy, usually provided by ATP
(adenosine triphosphate). Active transport serves to
maintain the normal balance of ions in cells, in particu-
lar ions of sodium and potassium, which play a vital
role in nerve and muscle cells. Because a molecule is
“pumped” across the membrane against its gradient
with the help of metabolic energy, it is referred to as
“active” transport.
The sodium–potassium “pump” that exchanges
sodium (Na+) for potassium (K+) across the plasma
membrane of animal cells is an example of the active
transport mechanism.
It is the carriage of a solute across a biological
membrane from low to high concentration that
requires the expenditure of metabolic energy.

adaptive radiation The process where a popula-
tion of plants or animals evolves into a number of dif-
ferent ones over time, usually as a response to
multiplying and living under different environmental
conditions. Subpopulations from the common ances-
tor develop as a response to adapting to the new envi-
ronmental conditions, and new species evolve from
this original parent stock.
Impressive rapid adaptive radiations have
occurred over time after mass extinctions caused by
cataclysmic episodes on the Earth. Plate tectonics, vol-
canism, and possible Earth-comet-asteroid collisions
all have wiped the landscape clean, allowing survivors
and new species to rapidly fill the voids of these new
adaptive zones.

address-message concept Refers to compounds
in which part of the molecule is required for
binding (address) and part for the biological action
(message).

adenosine 5’-triphosphate (ATP) Key NUCLEOTIDE
in energy-dependent cellular reactions, in combination
with Mg(II). The reaction: ATP + water Ô ADP +
phosphate is used to supply the necessary energy.
See alsoATP.

4 actinomorphic


Solanumspecies as an example of an actinomorphic flower from
Sierra Madre Oriental in Mexico.(Courtesy of Tim McCabe)

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