stimulated with Br to turn on disease resistance pathways, cleverly showing that each part of
these receptors is specific and can function when swapped.
Activation of the BRI receptor is thought to stimulate other protein kinases and phospha-
tases that help relay information to the nucleus where the BES1 transcriptional activator acts
to regulate gene expression. In this way the Br signal transduction pathway is similar to
other signaling pathways we have examined, with a final nuclear transcriptional output
required for the final biological response to Br. It is interesting to note that there are over
170 genes inArabidopsispredicted to encode LRR kinases that may function in plant
signal transduction. In addition, there are several other putative receptor-like kinases that
do not contain LRR domains, but also may function in signaling.
4.7 Conclusions
Even though plants do not have elaborate body plans and the number of specialized cells as
organs compared with animals, developmental programs are no less elaborate. A number of
crucial plant growth regulators or hormones are required for proper plant growth. We will
see in the next chapter how plant biotechnologists can alter hormone type and concentration
to manipulate cells in petri dishes, a requirement for plant genetic transformation.
LIFE BOX 4.1. NATASHA RAIKHEL
Natasha Raikhel, University Distinguished Professor, Ernst and Helen
Leibacher Endowed Chair, University of California Riverside
Natasha Raikhel
I originated from and grew up in the
Soviet Union. I immigrated with my
husband and first born son to Athens,
Georgia in 1978 (my second son was
born in Athens, Georgia) with a personal
fortune of only $25. I remember feeling
somewhat lost and wondered how I
could and would ever make the
language, scientific and social tran-
sitions required of me. I did not realize
at the time that I was lucky in many
ways and that fortune had favored me.
I knew only one American scientist
when I first arrived, but I encountered
many helpful people that were critical
to my survival. I also entered a social
context within academia that differed in
several important ways from the system
I left behind. The American academic
system is characterized by greater diver-
sity and openness of thought and a
healthy atmosphere of competition that
drives one to take intellectual risks and
achieve more. At its best, this environ-
ment also leads to a constant renewal
of possibility, a wealth of new ideas
and a rich milieu of thoughtful exchange
that fosters both collective and individ-
ual progress. In America, I found a
place where prestige and intellectual
and economic rewards were all reason-
able potential goals. Although I did not
106 PLANT DEVELOPMENT AND PHYSIOLOGY