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CHAPTER
8
Properties of Sensory
Receptors
OBJECTIVES
After studying this chapter, you should be able to:
■
Describe the classification of sensory receptors.
■
Name the types of sensory receptors found in the skin, and discuss their relation to
touch, cold, warmth, and pain.
■
Define generator potential.
■
Explain the essential elements of sensory coding.
INTRODUCTION
Information about the internal and external environment
activates the CNS via a variety of
sensory receptors.
These
receptors are transducers that convert various forms of energy
in the environment into action potentials in neurons. The
characteristics of these receptors, the way they generate
impulses in afferent neurons, and the general principles or
“laws” that apply to sensation are considered in this chapter.
Emphasis is placed on receptors mediating the sensation of
touch, and later chapters focus on other sensory processes.
We learn in elementary school that there are “five senses,” but
this dictum takes into account only some of the senses that
reach our consciousness. In addition, some sensory receptors
relay information that does not reach consciousness. For exam-
ple, the muscle spindles provide information about muscle
length, and other receptors provide information about arterial
blood pressure, the temperature of the blood in the head, and
the pH of the cerebrospinal fluid. The list of senses in Table 8–1
is somewhat simplified. The rods and cones, for example,
respond maximally to light of different wavelengths, and three
different types of cones are present, one for each of the three
primary colors. There are five different modalities of taste:
sweet, salt, sour, bitter, and umami. Sounds of different pitches
are heard primarily because different groups of hair cells in the
cochlea are activated maximally by sound waves of different
frequencies. Whether these various responses to light, taste,
and sound should be considered separate senses is a semantic
question that in the present context is largely academic.
SENSE RECEPTORS
& SENSE ORGANS
It is worth noting that the term
receptor
is used in physiology
to refer not only to sensory receptors but also, in a very differ-
ent sense, to proteins that bind neurotransmitters, hormones,
and other substances with great affinity and specificity as a
first step in initiating specific physiologic responses.
CLASSIFICATION OF
SENSORY RECEPTORS
Numerous attempts have been made to classify sensory recep-
tors, but none has been entirely successful. One classification
divides them into (1) teleceptors (“distance receivers”), which
are concerned with events at a distance; (2) exteroceptors,
which are concerned with the external environment near at
hand; (3) interoceptors, which are concerned with the internal