396 DAIRY CHEMISTRY AND BIOCHEMISTRY
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EXTERNAL Lactose Lactose Lactose
ENVIRONMENT
CELL WALL MEMBRANE I’MF
CYTOI’LASM
Laciore-P Lactose i Laclose J.
I
t
Glucose Lactose Glucose
Galactose-6-P p4-t ADI’ Glucose-6-P - Galactose-I-P GlUCoSe-I-P KT1’ t AUI’ + - Glucore.6.P LlDP
{C::::
6-Phosphogluconate
1
Fructose-6-P
J
Tagatose.6-P
k”, K ::,
$
t
co2
ADI’ t
$.
Ribulose-5-P
Tagatose.1.6.blP Fructose-1’6-hiP
Xylulose.5-P
p,
v
Acetyl-P
1
CuASH
Dthydroxyacetone-P -1 Glyceraldehyde-3-P t
:::,yp
1.3-Diphosphoglycerate
3-Phosphoglycerate 24 ACETY L-CoA
CnASH
?-Phosphoglycerate t
I
Tagatose
pathway
Phosphoenolpyruvate
A,.. ‘\I I’ Fyruvare 2
Lactate
Glycolytic pathway
ACETYLALDEHYDE NADH
K l\AIl’
Ethanol
Leloir Phosphoketolase
pathway pathway
Figure 10.12 Metabolism of lactose by lactic acid bacteria; many Lactobacillus species/strains
can not metabolize galactose (from Cogan and Hill, 1993).
Solubilizes of colloidal calcium phosphate and thereby affects cheese
texture; rapid acid production leads to a low level of calcium in the cheese
and a crumbly texture (e.g. Cheshire) and vice versa (e.g. Emmental).
Promotes syneresis and hence influences cheese composition.
Influences the activity of enzymes during ripening, and hence affects
cheese quality.