Colonization-related Trade-offs in Tropical Forests 193make possible some calculations of the role of
colonization-related trade-offs.
Finally, research on these trade-offs should go
beyond simply trait measurements and theory, to
other and stronger tests of the roles of the trade-
offs in real communities. Community-level field
studies of spatio-temporal variation in environ-
mental conditions, seed arrival, and successful
recruitment should be conducted to determine
the degree to which different mechanisms can
explain recruitment patterns. Field experiments
involving manipulations of seed arrival, seedling
recruitment, and environmental conditions could
provide even stronger tests of the mechanisms.
For time scales and spatial scales over which
such field studies and experiments are infeasible,
models parametrized from field data can provide
a useful tool to explore long-term, large-scale
dynamical implications of documented processes.
With this combination of tools, we should be
able to achieve a much better understanding of
colonization-related trade-offs in tropical forests in
the future.
ACKNOWLEDGMENTS
I thank Joe Wright, Steve Pacala, and two anony-
mous reviewers for helpful comments on this
chapter.
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