vertebral canals fall within the range of modern humans.Whereas some
may debate interpretation of vertebral arch size,it is evident that the
thoracic cage perfectly matches the modern human condition.
At the other end of the respiratory tract,a modern humanlike exter-
nal nose also makes its appearance by about 1.5 million years ago.First
described in detail by Franciscus and Trinkaus (1988),the bony archi-
tecture forming the structure of the human external nose is virtually
identical in australopithecines and chimpanzees (and other apes),but is
distinctly humanlike in fossils such as ER-3733 and others identified as
Homo erectusor early Homo sapiens(figure 14.4).In apes and aus-
tralopithecines,the nasal bones are flat (from side to side and top to
bottom) and the upper nasal margins are even with or sunken below the
transverse plane of the orbits.In living humans,the nasal bones are
curved and arched forward,and the upper nasal area projects away
from the face forming a true external nose.Franciscus and Trinkaus
(1988:524–525) emphasized important increases in “effectiveness in
humidification and temperature modification of inspired air”these
anatomical modifications signal,but development of the external nose
also has implications for speech sounds,since some consonants (e.g.,/n/)
are produced by nasal resonance.Apes apparently lack the ability to
produce consonants (Savage-Rumbaugh and Lewin 1994),which may in
part relate to their lack of an external nose.However,with the earliest
227 Fossil Evidence for the Origin of Speech Sounds
Figure 14.4
Views of nasal areas of a chimpanzee (left) and an early Homo (right).In the chimpanzee
the nasal bones are depressed and level with the sides of the face.In the early Homo(cast
of ER-3733) the nasal bones are arched and project away from the face,forming an exter-
nal nose.
Fig.14.4