variations grouped around that pattern.The accomplishments of a winter
wren are of an altogether different magnitude.Each male has a reper-
toire of maybe ten song types and the winter wren is only a beginner as
wrens go;other wren species have repertoires numbered in the hundreds
(Kroodsma and Verner 1978).So this is the first bird-ape contrast.Some
songbirds with learned songs have individual repertoires that are huge;
repertoires of monkeys and apes are strictly limited.
If we examine the way in which large birdsong repertoires develop,we
find an ontogenetic principle operating that is either feeble or simply
lacking in nonhuman primates.Vocalizations of monkeys and apes are
innate.Songbirds,with their learned songs,have a developmental strat-
egy with all the hallmarks of a truly creative process.As far as I know,
phonocoding with this degree of richness has never been recorded in any
animal with innate vocalizations.In a classic manifestation of phonolog-
ical syntax,wrens and other songbirds display a remarkable ability to
rearrange learned phrases,seemingly doing so almost endlessly in some
species.Many different sequences are created,generating,in effect,a
kind of animal music.The sequences are not random but orderly,orga-
nized by definable rules and structured in such a way as to yield many
stable,repeatable,distinctive patterns,the precise number varying from
species to species and bird to bird.Together with whale songs,also
learned (Payne,Tyack,and Payne 1983;Payne,this volume),these bird-
songs are an obvious place to look for insights into what underlying aes-
thetic principles,if any,are shared between animal and human music.
I draw two modest conclusions from this overview of animal commu-
nication,language and music.From a reductionistic point of view,a
convenient basis for animal-human comparisons is provided by the
realization that the potential for phonocoding is a critical requisite for
the emergence not only of speech and language but also of music.I
suggest that we can already see a version of such a process in operation,
albeit in primordial form,in some learned songs of animals,especially
songbirds.An obvious next step would be to analyze phonocoding rules
that birds use when they sing.Is it possible that they conform to our own
compositional rules? A minimalist definition of music,at least of the
Western,tonal variety,might be couched in terms of notes with specific
pitches,intervals,and distinctive timbres combined into phrases that are
repeated with additions and deletions,assembled into series with a par-
ticular meter and rhythm,and so constituting a song or melody.One
approach to the human-animal comparison would be to see whether any
animal sounds conform to similar taxonomic criteria,all of which are
potentially studiable in animals.Do the rules vary among species in rela-
tion to repertoire size? Are correlations seen among lifestyle,tempera-
ment,and song tempo in different bird species? It is my conviction that
45 Origins of Music and Speech
MUS3 9/14/99 11:58 AM Page 45