with sporangia at their tips (Fig. 3). Their relationships are obscure, perhaps
with the pteridosperms, but their leaves resemble microphylls such as possibly
those of progymnosperms (Topic Q4). If they are related to the progymno-
sperms it suggests that Cordaitales and conifers may have developed the seed
habit independently from pteridosperms and other groups.
There were several other groups of seed plants found only as fossils that had
died out by the end of the Cretaceous period. Five groups of seed plants
survive. Of these, the conifers have a fossil history extending right back to the
Carboniferous, the cycads to the Triassic and the Ginkgoales as far as the
Permian period. The Gnetales and the angiosperms are more recent, probably
appearing first in the Jurassic period.
R1 – Early seed plants 295
Microsporangia
1 mm
Fig. 3. Cone-like male reproductive structure of the Cordaitales.