- According to the sliding filament theory, actin and
myosin slide past each other to produce sarcomere
shortening. Ca2+is released in the sarcomere in response
to an action potential that exposes myosin cross-bridge
binding sites on actin. Myosin cross bridges bind to
actin and pull actin filaments closer to the center of each
sarcomere producing force and stiffness within the
skeletal muscle (Lorenz and Campello, 2001).
•Force production by a skeletal muscle can be volun-
tarily graded; however, the muscle fibers innervated
by one motor neuron (i.e., a motor unit) act in an all
or none fashion. As more motor units are recruited in
a particular skeletal muscle, the muscle produces
more force (Lorenz and Campello, 2001).
SKELETAL MUSCLE FIBER TYPES
- There are several different types of muscle fiber based
on structure and function. These include Type I, Type
IIA, and Type IIB.
•Type I muscle fibers (slow twitch oxidative fibers)
have high mitochondria content and a rich blood
supply. They are difficult to fatigue but do not provide
high amounts of muscle force or tension. Type I fibers
act in an aerobic capacity during activity.
•Type IIA muscle fibers (fast twitch oxidative-
glycolytic fibers) also have high mitochondria content
and are moderately capable of performing aerobic and
anaerobic activities.
•Type IIB muscle fibers (fast twitch glycolytic fibers)
have sparse mitochondria content and blood supply.
They fatigue easily but are capable of producing
higher force and tension. Type IIB fibers act in an
anaerobic capacity during activity. - All muscle fibers within a motor unit have the same
metabolic characteristics (fiber type); however, skele-
tal muscles can be composed of different types of
muscle fibers based on the metabolic demand of the
muscle (Lorenz and Campello, 2001).
•Muscle fiber type composition in human skeletal
muscle is genetically determined (Lorenz and
Campello, 2001). There is little information regarding
muscle fiber type transformations in response to train-
ing/exercise. - Heavy resistance training caused increased type IIA
and decreased type IIB fibers, while type I fiber com-
position in human skeletal muscle was unchanged
(Adams et al, 1993). - Structural and genetic characteristics of muscle fiber
types have been modulated with fiber-specific stimu-
lation in vitro (Liu et al, 2001); however, it is
unknown whether such changes occur in all muscle
fiber types or if the transformation will be sustained
over time in vivo.- Since muscle fiber composition is genetically deter-
mined, athletes may participate in sports or activities
that involve muscle contractions that are more natural.
Whether a distance runner can train to be a successful
power lifter or vice versa is an issue that has not yet
been clearly elucidated.
- Since muscle fiber composition is genetically deter-
TYPES OF SKELETAL MUSCLE CONTRACTION
•Skeletal muscle can produce joint movements through
concentric and eccentric movements. Skeletal muscle
contractions produce muscle tension and control body
and joint movement.
- Concentric contractions describe a movement that
involves shortening of a muscle against a load,
whereas eccentric contractions involve controlled
lengthening of a muscle against a load. - Eccentric muscle contractions produce greater muscle
force and more myofibrillar disruption than concen-
tric exercise (Gibala et al, 1995). - Eccentric muscle contractions (often referred to as
negativesin weightlifting) are more effective in pro-
ducing strength gains and hypertrophy than concen-
tric contractions (Higbie et al, 1996); however, both
eccentric (negative) and concentric (positive) contrac-
tions elicit gains in skeletal muscle strength and size
(Higbie et al, 1996). - Isometric muscle contractions produce muscle tension
without joint movement. For example, pushing
against a wall or contracting the quadriceps muscle
while holding the knee motionless at a particular point
in the knee range of motion. - Isotonic muscle contractions produce muscle tension
and joint movement against a constant load where rate
of movement is variable. For example, a dumbbell
curl is a contraction against a constant load that can be
voluntarily moved at a self-selected rate. This is the
most typical contraction in weightlifting. - Isokinetic muscle contractions involve a constant rate
of joint displacement that is maintained by varying
amounts of resistance based on muscle effort. This is
uncommon in weightlifting or athletic settings.
Isokinetic exercise requires expensive machinery and
is usually most applicable in the rehabilitation setting. - Isotonic and isokinetic muscle movements can be per-
formed through concentric or eccentric muscle con-
tractions.
MUSCLE RESPONSE TO RESISTANCE TRAINING
•Improvements in muscle strength, power, or
endurance are best achieved by overloading the
muscle(s) being trained.
544 SECTION 6 • SPORTS-SPECIFIC CONSIDERATIONS