B_T_2015_02_

(Michael S) #1

  1. octaploid Triticale

  2. hexaploid Triticale


(c) Wheat


• Common wheat (Triticum aestivum) is an important
example of natural allopolyploid.


•    Common wheat is hexaploid, i.e., 2n = 6x = 42 with
three genomes, A, B and D, i.e., AABBDD.
• These three genomes are derived from three different
diploid species, such as :


  • Triticum aegilopoides – AA (2n = 14).

  • Aegilops speltoides – BB (2n = 14).

  • Aegilops squarossa – DD (2n = 14).
    • By crossing first two diploid species, Triticum
    dicoccoides, 2 n = 4x = 28, i.e., tetraploid – AABB,
    is produced, which on crossing with third species
    gives rise to Triticum aestivum, 2 n = 6x = 42, i.e.,
    hexaploid – AABBDD.


(iii) autoallopolyploidy or segmental poly-
ploidy
• This is intermediate between the first two types of
polyploidy.
• Wheat is nowadays considered to be an example of
segmental polyploidy, rather than allopolyploidy, as
earlier thought because three genomes in wheat A,
B and D, are not fully distinct from each other, and
have some portions common in them (i.e., only some
segments are homologous).
• Hence A, B and D show homoeologous relationship
(not homologous), so pairing can occur upto some
extent.
• Such polyploids where different genomes show
homoeologous relationship are called autoallopolyploids
or segmental polyploids.

Intext Practice Questions



  1. Explain the different types of polyploidy.

  2. Fill in the blanks with appropriate words/phrases :
    (a) Organisms having genetic constitution, 2n – 2, are called __.
    (b) Aneuploidy occurs due to __ during meiosis.
    (c) Organisms with genetic constitution, 2n – 1 – 1, are called __.


role in evolution
• One of the most frequent cause of evolution
is a change in the order of genes as a result of
chromosomal aberrations.
• The majority of plant species originate from an abrupt
and rapid change in nature, and aneuploidy and

polyploidy are the prime sources of variation.
• In the animal kingdom polyploidy is not so important.
Among vertebrates, different species of fishes have
a different number of chromosomes. Amphibians
are generally characterized by a specific number for
each family.
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