FIGURE 13.16. The family tree of the dogs, showing the variation from weasel-like Hesperocyon to the huge
hyena-like bone-crushing borophagine dogs. (Based on information supplied by Xiaoming Wang; drawing
by Carl Buell)
Eocene Oligocene Miocene Plio Pl
Million
Years
Hesperocyon - Paraehydrocyon
Mesocyon - Enhydrocyon
Ectopocynus
Osbornodon
Otarocyon
Cynarctoides
Phlaocyon
Cormocyon
Desmocyon
Cynarctus
Tomarctus
Aelurodon + Strobodon
Paratomarctus
Carpocyon
Epicyon
Osteoborus
Borophagus
Leptocyon
Foxes
Eucyon
S. Amer.
Canines
Canines
Borophagus
Hesperocyon Canis
Tomarctus
Enhydrocyon
Borophagus
Aelurodon
40 36 32 28 24 20 16 12 8 4 0
Hesperocyon