Flora Unveiled

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Crop Domestication and Gender j 37

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Of all the differences between wild and domesticated cereals, two are regarded as
most critical for domestication. The first is the loss of seed dormancy— the inability of
certain seeds to germinate when planted in moist soil unless given additional environ-
mental stimuli, such as a cold treatment. Because the loss of seed dormancy trait is invis-
ible, it does not show up in the archeological record. Such a change may have begun
either during the phase of ecosystem management prior to the adoption of a sedentary
lifestyle, or later, in small garden plots planted near permanent settlements during the

(a)
(b)

(c)
palea
stigma

lemma

carpel

anther + lament

Figure 3.3 Three views of the oat flower. A. An oat spikelet contains a pair of florets. B. Oat flower
dissected from a single floret. C. Diagram of oat floret. The ovary, topped by two feathery stigmas, is
surrounded by three stamens. The reproductive structures are enclosed in two modified scales, the
larger lemma with its bristle- like extension, the awn, and the smaller palea (missing in B). During
selfing, pollen released from the anthers is deposited on the stigmas of the same flower before the
opening of the floret.
B is from: http:// http://www.biology.iastate.edu/ Courses/ 202L/ New%20Site%20S05/ 27AngioReprod/ Plantrepro.htm).
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