Biology Today - May 2018

(Rick Simeone) #1

  • The process of development of new organisms through the formation and fusion of gametes is called sexual reproduction.

  • In angiosperms, the organs specialised to perform sexual reproduction are flowers.

  • Flowers are modified condensed reproductive shoots.

  • A typical flower has a broad base called thalamus over which four whorls of floral leaves, i.e., sepals (calyx), petals (corolla),
    stamens (androecium) and carpels (gynoecium) are borne.

  • Stamens and carpels represent male and female reproductive structures of flowers respectively. They are called essential floral organs.

  • Sepals and petals are called non-essential floral organs because they have only supportive role.


STAMEN-THE MALE REPRODUCTIVE ORGAN



  • Stamen consists of two parts-filament and anther.

  • Filament is long and slender stalk attached proximally to thalamus, petal or tepal. It bears an anther distally.

  • Anther is bilobed, knob-like fertile part of the stamen. The two anther lobes separated by a deep groove in the anterior side
    and attached to each other by a sterile parenchymatous tissue called connective on the posterior side.

  • Each anther lobe has two chambers which posses pollen sacs or microsporangia.

  • A bilobed anther is tetrasporangiate.

  • A microsporangium or pollen sac is a cylindrical sac which appears circular in transverse section.

  • It consists of two parts : outer wall and central homogeneous sporogenous tissue.

  • The outer wall has four types of layers epidermis, endothecium, 1-3 middle layers and tapetum.

  • The outer three layers protect the young anther and take part in the mechanism of dehiscence in the ripe anther.

  • The endothecium is also called fibrous layer due to the presence of fibrous thickenings.

  • The tapetal cells enlarge and become filled with dense protoplasmic content as well as nutrients.

  • Two types of tapetum are present amoeboid and secretory.


Functions of Tapetum


  • It provides nourishment to the developing microspore mother cells and pollen grains either by forming a plasmodium (amoeboid
    type) or through diffusion (secretory type).

  • It produces lipid rich Ubisch granules containing sporopollenin for exine formation.

  • It secretes enzymes like callase responsible for degradation of callose wall around pollen tetrad.


Class XII


Sexual Reproduction in Flowering Plants


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