118
relationships with a 54 kb region on rice chromosome 12. A total of fi ve markers
were mapped within a genetic interval of 0.3 cm and 400 kb BAC contigs were
established on both sides of the gene to lay the foundation for the map-based cloning
of SKr. Two SSR markers completely linked to SKr were used to evaluate a collec-
tion of crossable wheat progenies originating from a primary triticale breeding
programme (Alfares et al. 2009 ). The marker cfb306 , which is closely linked to SKr ,
was found to be an effi cient marker for the int roduction of crossability into wheat
germplasm through marker-assisted selection. In work performed by Mishina et al.
( 2009 ) QTL mapping was carried out using the F 7 population derived from a cross
between Chinese Spring (CS) and a CS/Cheyenne 5B chromosome substitution
line. In this population, a major QTL region controlling crossability with rye was
detected on a locus closely linked to the SSR marker Xgwm443 on the short arm of
chromosome 5B, which was assumed to be the Sk r locus.
Acknowledgements This work was supported by the Hungarian National Scientifi c Research
Fund (OTKA K 104382) and by the “Wheat and Barley Legacy for Breeding Improvement”—
WHEALBI EU FP7 project. The technical assistance of Edina Türkösi is gratefully acknowledged.
Thanks are due to Barbara Hooper for revising the manuscript linguistically.
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