The New Neotropical Companion

(Elliott) #1

Peru. Two Amazonian tributary rivers, the Negro and the
Madeira, rank as the fifth- and sixth- largest rivers in the
world with regard to annual discharge. By comparison,
the Mississippi River ranks about 10th and has only about
1/12 the annual discharge of the Amazon.
To the surprise of many, the Amazon River originally
flowed in the opposite direction, draining into the Pacific
Ocean near what is today the port city of Guayaquil,
Ecuador. The river changed to its present west- to- east
course as recently as 10– 15 million years ago, when the
Andean uplift profoundly altered the river’s course, as
well as patterns of biogeography, creating the Amazon
Basin. Initially the uplift of the Andes created a gigantic
lake, bordered on the west by the newly arisen mountain
chain and to the east by the extensive Guiana and


Brazilian Shields. The Amazon finally made its way to
the Atlantic during the Pleistocene, cutting through its
eastern barrier in the vicinity of Obidos, Brazil. Many
widespread trees were probably dispersed eastward by
the altered course of the river water.
The Amazon Basin, drained by the Amazon River
and its gigantic tributaries, covers an area of about 6.92
million km^2 (about 2.67 million mi^2 ) essentially 40% of
the total area of South America. Approximately 1,100
tributaries service the main river, and some of them,
like the Negro, Napo, Madeira, Tapajós, Tocantins, and
Xingu, rank as major rivers.
Amazon tributaries vary in color from cloudy mocha
to clear black depending upon where they originate and
their geological and chemical properties (chapter 6).

COLOMBIA

Negro
Branco

Tapajos

Japura

Jurua

Putumayo

Marañón

Ucayali

Lima
PERU

Pacific
Ocean

BOLIVIA

Santa Cruz

Cusco La Paz

Rio Branco

Madre
de Dios

Pucallpa

Iquitos

Cochabamba

Guapore

Mamore

Beni

Pôrto
Velho
Juruena Xingu

Machado

Teles Pires

Araguai

a

BRAZIL

São Paulo

Rio de
Janeiro

Brasilia

Belem

Macapá

Tocantins

Purus Madeira

VENEZUELA

Bogota

Quito

Caracas

Georgetown
GUYANA

Atlantic
Orinoco Ocean

Amazon
Santarém
Manaus

ECUADOR

Figure 12– 2. Map of the Amazon River and its major tributaries. Reprinted with permission from Kricher, John. Tropical Ecology.
Princeton, NJ: Princeton University Press, 2011.


208 chapter 12 cruising the rivers to the sea

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