468 MHR • Unit 5 Population Dynamics
Similar methods are used in studies of aquatic
ecosystems to estimate the size of various
populations. Often, samples of a known volume of
water are collected. The water is then passed through
a net or sieve, and the number of organisms in each
sample is counted. As with quadrats, the average
density (number of individuals per unit volume)
can be used to estimate the size of the population
contained in the entire body of water.
When using any sampling technique, it is
important to take randomsamples — that is, samples
in which all individuals in the population have an
equal chance of being represented. To have the best
chance of taking such a sample, it is important to
know something about how the individuals in the
population are distributed — in other words, to
understand the population’s dispersion. In the
MiniLab on the next page, you will practise one
technique used for estimating populations of
mobile species.
Population Dispersion
Imagine walking along a transect and making a note
on your data sheet each time you see a member of
the population in which you are interested. Do you
think you would be counting individuals at regular
intervals — perhaps one every 10 m? Or do you
think it more likely that you would come across
several individuals at once, and then none, until
you reach another group? This, of course, depends
on how the members of the population are dispersed.
To simplify discussion, ecologists tend to
recognize three general patterns of dispersion:
uniform, random, and clumped (see Figure 14.5).
It is important to realize, however, that there is
actually a continuum of dispersion patterns. Many
species typically display patterns that fall between
two of these three types. Factors important in
determining dispersion include the distribution of
Ecologists are often more interested in knowing the relative
densities of two or more populations (that is, whether one
is more or less dense than another) rather than the actual
or absolutedensity of each one. Relative densities can be
determined by looking for signs of activity along transects.
For example, the relative density of bears in an area is often
estimated by looking for footprints, claw marks on trees,
clumps of hair caught on bushes, and piles of droppings
(usually called “scats” by biologists). Since it is difficult to
distinguish between grizzly (Ursus horribilis) and black
bears (U. americanus) using some of these signs, samples
of hair or scats are brought back to the lab for DNA analysis.
BIO FACT
Figure 14.4When sampling the density of certain plants in
a quadrat, it is often too difficult to count the number of
individuals of a species. In this case, the proportion of the
quadrat covered or shaded by one or more plant species is
recorded. The area sampled is usually square or circular.