Biological Oceanography

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First nauplii (N1) were obviously undersampled, possibly hatching and staying too
close to the bottom to catch. Recruitment to N2, mostly determined by egg input,
clearly had strong variability in time. This is usually due in part to variability in
female abundance and fecundity, but intense and variable mortality of eggs and N1
hatchlings is also important. Whatever the source of variability, its pattern both
persists and evolves as the animals proceed through their developmental stages.
Landry picked out somewhat arbitrary “cohorts” and shaded them alternately in his
time-series representation (Fig. 7.11). Carbon contents of each stage in each cohort
were determined by CHN (carbon, hydrogen, nitrogen) analyzer. The date at which a

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