Chesson, P. (2000) Mechanisms of maintenance of species
diversity.Annual Review of Ecology and Systematics, 31 ,
343–66.
Clark,J.S.,Dietze,M.,Chakraborty,S.,et al. (2007) Resolving
the biodiversity paradox.Ecology Letters, 10 , 647–62.
Condit, R., Pitman, N., Leigh Jr, E.G.,et al. (2002) Beta
diversity in tropical forest trees.Science, 295 , 666–9.
Connor, E.F. and McCoy, E.D. (1979) The statistics and
biology of the species-area relationship.The American
Naturalist, 113 , 791–833.
Cornell, H.V. (1993) Unsaturated patterns in species as-
semblages: the role of regional processes in setting local
species richness. InSpecies Diversity in Ecological Com-
munities: Historical and Geographical Perspectives(eds R.E.
Ricklefs and D. Schluter ), pp. 243–52. University of
Chicago Press, Chicago, IL.
Damschen, E.I., Haddad, N.M., Orrock, J.L.,et al. (2006)
Corridors increase plant species richness at large scales.
Science, 313 , 1284–6.
Diamond, J.M. (1975) Assembly of species communities.
InEcology and Evolution of Communities(eds M.L. Cody
and J.M. Diamond), pp. 342–444. Harvard University
Press, Cambridge, MA.
Drakare, S., Lennon, J.J. and Hillebrand, H. (2006) The
imprint of the geographical, evolutionary and ecologi-
cal context on species-area relationships.Ecology Letters,
9 , 215–27.
Etienne, R.S., Alonso, D. and McKane, A.J. (2007) The
zero-sum assumption in neutral biodiversity theory.
Journal of Theoretical Biology, 248 , 522–36.
Forbes, A.E. and Chase, J.M. (2002) The role of habitat
connectivity and landscape geometry in experimental
zooplankton metacommunities.Oikos, 96 , 433–40.
Fukami, T. (2004) Community assembly along a species pool
gradient: implications for multiple-scale patterns of spe-
cies diversity.Population Ecology, 46 , 137–47.
Gause, G.F. (1934)The Struggle for Existence. Williams &
Wilkins, Baltimore, MD.
Gilbert, F., Gonzalez, A. and Evans-Freke, I. (1998) Corri-
dors maintain species richness in the fragmented land-
scapes of a microecosystem.Proceedings of the Royal
Society of London B, 265 , 577–82.
Gotelli N.J. and Ellison, A.M. (2006) Food-web models
predict species abundances in response to habitat
change.PLoS Biology, 4 , e324.
Gravel, D., Canham, C.D., Beaudet, M. and Messier, C.
(2006) Reconciling niche and neutrality: the continuum
hypothesis.Ecology Letters, 9 ,399–409.
Hanski, I. and Gyllenberg, M. (1997) Uniting two general
patterns in the distribution of species.Science, 275 ,
397–400.
Harrison, S. (1997) How natural habitat patchiness affects
the distribution of diversity in Californian serpentine
chaparral.Ecology, 78 , 1898–906.
Harrison, S. (1999) Local and regional diversity in a patchy
landscape: native, alien and endemic herbs on serpen-
tine soils.Ecology, 80 , 70–80.
Harrison, S., Davies, K.F., Safford, H.D. and Viers, J.H.
(2006a). Beta diversity and the scale-dependence of the
productivity-diversity relationship: a test in the Califor-
nian serpentine flora.Journal of Ecology, 94 , 110–17.
Harrison, S., Safford, H.D., Grace, J.B.,et al. (2006b). Re-
gional and local species richness in an insular environ-
ment: serpentine plants in California. Ecological
Monographs, 76 , 41–56.
Hastings, A. (1980) Disturbance, coexistence, history, and
competition for space.Theoretical Population Biology, 18 ,
363–73.
Holt, R.D. (1993) Ecology at the mesoscale: the influence of
regional processes on local communities. InSpecies Di-
versity in Ecological Communities(eds R.E. Ricklefs and
D. Schluter), pp. 77–88. University of Chicago Press,
Chicago, IL.
Holt,R.D.andHoopes,M.F.(2005)Foodwebdynamicsina
metacommunity context: modules and beyond. InMeta-
communities: Spatial Dynamics and Ecological Communities
(eds M. Holyoak, M. Leibold and R. Holt), pp. 68–93.
University of Chicago Press, Chicago, IL.
Holt, R.D., Lawton, J.H., Polis, G.A. and Martinez, N.D.
(1999) Trophic rank and the species-area relationship.
Ecology, 80 , 1495–505.
Holyoak, M. and Loreau, M. (2006) Reconciling empirical
ecology with neutral community models.Ecology, 87 ,
1370–7.
Holyoak,M.,Leibold,M.A.andHolt,R.D.,eds.(2005)Meta-
communities: Spatial Dynamics and Ecological Communities.
University of Chicago Press, Chicago, IL.
Horn, H. and MacArthur, R.H. (1972) Competition among
fugitive species in a harlequin environment.Ecology, 53 ,
749–52.
Hoyle, M. and Gilbert, F. (2004) Species richness of moss
landscapes unaffected by short-term fragmentation.
Oikos, 105 , 359–67.
Hubbell, S.P. (2001)The Unified Neutral Theory of Biodiver-
sity and Biogeography. Princeton University Press, Prin-
ceton, NJ.
Huffaker, C.B. (1958) Experimental studies on predation:
dispersion factors and predator-prey oscillations.Hil-
gardia, 27 ,343–83.
Hugueny, B., Cornell, H.V. and Harrison, S. (2007) Meta-
community models predict the local-regional richness
relationship in a natural system.Ecology, 88 , 1696–706.
REFERENCES 213