MacArthur, R.H. and Wilson, E.O. (1967)Theory of Island
Biogeography. Princeton University Press, Princeton, NJ.
Mouillot, D. (2007) Niche-assembly vs. dispersal-assem-
bly rules in coastal fish metacommunities: implications
for management of biodiversity in brackish lagoons.
Journal of Applied Ecology, 44 , 760–7.
Mouquet, N. and Loreau, M. (2003) Community patterns
in source-sink metacommunities.The American Natural-
ist, 162 , 544–57.
Nee, S. and May, R.M. (1992) Dynamics of metapopula-
tions: habitat destruction and competitive coexistence.
Journal of Animal Ecology, 61 , 37–40.
Norberg, J., Swaney, D.P., Dushoff, J.,et al. (2001) Pheno-
typic diversity and ecosystem functioning in changing
environments: a theoretical framework.Proceedings of
the National Academy of Sciences of the United States of
America, 98 , 11376–81.
O ̈berg, S., Ekbom, B. and Bommarco, R. (2007) Influence
of habitat type and surrounding landscape on spider
diversity in Swedish agroecosystems.Agriculture Eco-
systems and Environment, 122 , 211–19.
Odling-Smee, F.J., Laland, K.N. and Feldman, M.W. (2003)
Niche Construction: The Neglected Process in Evolution.
Princeton University Press, Princeton, NJ.
Oksanen, T. (1990) Exploitation ecosystems in heteroge-
nous habitat complexes.Evolutionary Ecology, 4 , 220–34.
O ̈stman, O ̈., Ekbom, B. and Bengtsson, J. (2001) Landscape
heterogeneity and farming practice influence biological
control.Basic and Applied Ecology, 2 , 365–71.
O ̈stman, O ̈., Ekbom, B. and Bengtsson, J. (2003) Yield
increase attributable to aphid predation by ground-liv-
ing natural enemies in spring barley in Sweden.Ecologi-
cal Economics, 45 , 149–58.
Palumbi, S.R. (2001) Humans as the world’s greatest evo-
lutionary force.Science, 293 , 1786–90.
Peterson, D.L. and Parker, V.T., eds. (1998)Ecological Scale:
Theory and Applications. Columbia University Press,
New York, NY.
Polis, G.A., Anderson, W.B. and Holt, R.D. (1997) Toward
an integration of landscape and food web ecology: the
dynamics of spatially subsidized food webs.Annual
Review of Ecology and Systematics, 28 , 289–316.
Ricklefs, R.E. and Schluter, D., eds. (1993)Species Diversity
in Ecological Communities: Historical and Geographical Per-
spectives. University of Chicago Press, Chicago, IL.
Roschewitz,I.,Gabriel,D.,Tscharntke,T.andThies,C.(2005)
The effects of landscape complexity on arable weed species
diversity in organic and conventional farming.Journal of
Applied Ecology, 42 , 873–82.
Rundlo ̈f, M. (2007) Biodiversity in agricultural landscapes:
landscape and scale-dependent effects of organic farm-
ing. PhD thesis, Lund University, Sweden.
Rundlo ̈f, M. and Smith, H.G. (2006) The effect of organic
farming on butterfly diversity depends on landscape
context. Journal of Applied Ecology, 43 , 1121–7.
Rundlo ̈f, M., Bengtsson, J. and Smith, H.G. (2008) Local
and landscape effects of organic farming on butterfly
species richness and abundance.Journal of Applied Ecol-
ogy, 45 , 813–20.
Schmidt, M.H. and Tscharntke, T. (2005) The role of perennial
habitats for Central European farmland spiders.Agricul-
ture Ecosystems and Environment, 105 , 235–42.
Schoener, T.W. (1976) The species-area relation within
archipelagos: models and evidence from island land
birds. InProceedings of the 16th International Ornithologi-
cal Congress Canberra, 1974(eds H.J. Firth and J.H. Ca-
laby), pp. 629–42. Australian Academy of Science,
Canberra, ACT.
Steffan-Dewenter, I., Mu ̈nzenberg, U., Bu ̈rger, C.,et al.
(2002) Scale-dependent effects of landscape context on
three pollinator guilds.Ecology, 83 , 1421–32.
Thies, C. and Tscharntke, T. (1999) Landscape structure
and biological control in agroecosystems.Science, 285 ,
893–5.
Tremlova, K. and Mu ̈nzbergova, Z. (2007) Importance of
species traits for species distribution in fragmented
landscapes.Ecology, 88 , 965–77.
Tscharntke, T., Klein, A.M., Kruess, A.,et al. (2005) Land-
scape perspectives on agricultural intensification and
biodiversity ecosystem service management.Ecology
Letters, 8 , 857–74.
van de Koppel, J., Bardgett, R., Bengtsson, J.,et al. (2005)
Effects of spatial scale on trophic interactions.Ecosys-
tems, 8 , 801–7.
Vandermeer, J. and Perfecto, I. (2007) The agricultural
matrix and a future paradigm for conservation.Conser-
vation Biology, 21 ,274–7.
Vitousek, P.M., Mooney, H.A., Lubchenco, J. and Melillo,
J.M. (1997) Human domination of earth’s ecosystems.
Science, 277 , 494–9.
Weibull, A.C. and O ̈stman, O ̈. (2003) Species composition
in agroecosystems: the effect of landscape, habitat and
farm management.Basic and Applied Ecology, 4 , 349–61.
Weibull, A.C., Bengtsson, J. and Nohlgren, E. (2000)
Diversity of butterflies in the agricultural landscape:
the role of farming system and landscape heterogeneity.
Ecography, 23 , 743–50.
Weibull, A.C., O ̈stman, O ̈. and Granqvist, A ̊. (2003)
Species richness in agroecosystems: the effect of land-
scape, habitat and farm management.Biodiversity and
Conservation, 12 , 1335–55.
Worster, D. (1994)Nature’s Economy: A History of Ecolog-
ical Ideas, 2nd edn. Cambridge University Press, Cam-
bridge.REFERENCES 227