b
S1
S2
S3
S4
K D
D
S
K
a
KeystoneNon-keystoneCompetitive dominantCompetitive subordinateNutrient most neededNutrientStrong nutrient uptakeWeak nutrient uptake
NK
Plate 2
Keystone consumer in (a) a small module and (b) a complex
food web. See page 39.