other, or moving past each other laterally. Relatively
few islands are ancient biogeographically; some
have experienced a complex history of both lateral
movements and alternating emergence and submer-
gence, with profound importance to the resulting
biogeography, not just of the particular island in
question, but also of the wider region (Keast and
Miller 1996).
Plate tectonic processes give rise to islands by
three main means. First, the breaking away of
pieces of continent by sea-floor spreading has
carried New Zealand, Madagascar, and a few other
ancient islands off into isolation (Box 2.2). Second,
in connection with plate boundaries, volcanic
islands may arise to form an archipelago of islands,
as in the case of the Greater and Lesser Sundas—the
many islands of the Indonesian region. Third, vol-
canic islands may arise from hotspots (e.g. the
Hawaiian islands) and certain parts of mid-ocean
ridges. Hawaii is of hotspot origin, and Iceland is
MODES OF ORIGIN 13
Trench
Sea level
Asthenosphere
Trench
(a)
mantle
Volcanic arc
Convergent
plate
boundary
Divergent
plate boundary
(mid-ocean ridge)
Convergent
plate boundary
(b)
Upper
Eurasian
plate
Pacific
plate Cocos
Nazca
plate
Antarctic plate
South
Eurasian
plate
African
plate
North
American
plate
90 °N
90 °E 120 °E 150°E 180 ° 150 °W 120°W90°W60°W30°W30 0 ° °E60°E90°E
60 °N
30 °N
30 °S
60 °S
75 °S
0 °
Indo-
Aust.
plate
American
plate
Continental
crust
Indo-Aust.
plate
Upper mantle
Oceanic crust
Oceanic
crust
Figure 2.1Model and map of crustal plates. (a) The basic plate tectonics model, exemplified by the South Pacific. New oceanic crust and upper
mantle are created along the divergent plate boundary and spread east and west from here. The eastward moving plate (the Nazca plate) is
subducted beneath the continental lithosphere of South America. The westward movement of the much larger Pacific plate ends with subduction
beneath the oceanic lithosphere of the Indo-Australian plate. At both trenches, the subducted plate begins to melt when it reaches the
asthenosphere. (b) The major plates of the world. Divergent plate boundaries (mid-ocean ridges), at which plates move apart, are represented by
parallel lines. Convergent plate boundaries (mostly marked by trenches) are represented by lines with teeth along one side: the teeth point from
the downward moving plate towards the overriding or buoyant plate. Transverse plate boundaries are shown by solid lines. Broken lines indicate
boundaries of an uncertain nature. (Redrawn from Nunn 1994, Fig. 2.4.)