species in a variable environment: the problem of
reserve design in the Northern Territory, Australia.
Global Ecology and Biogeography Letters, 2 , 1–10.
Worthen, W. B. (1996). Community composition and
nested-subset analyses: basic descriptors for commu-
nity ecology. Oikos, 76 , 417–426.
Wragg, G. M. (1995). The fossil birds of Henderson Island,
Pitcairn Group: natural turnover and human impact, a
synopsis.Biological Journal of the Linnean Society, 56 ,
405–414.
Wright, D. H. (1983). Species–energy theory: an extension
of species–area theory. Oikos, 41 , 496–506.
Wright, D. H., Currie, D. J., and Maurer, B. A. (1993).
Energy supply and patterns of species richness on local
and regional scales. In Species diversity in ecological
communities: historical and geographical perspectives(ed.
R. Ricklefs and D. Schluter), pp. 66–74. University of
Chicago Press, Chicago.
Wright, D. H., Patterson, B. D., Mikkelson, G. M., Cutler,
A., and Atmar, W. (1998). A comparative analysis of
nested subset patterns of species composition.
Oecologia, 113 , 1–20.
Wright, D. H. and Reeves, J. J. (1992). On the meaning and
measurement of nestedness of species assemblages.
Oecologia, 92 , 416–428.
Wright, S. (1932). The roles of mutation, inbreeding,
crossbreeding, and selection in evolution. Proceedings
of the XI International Congress of Genetics 1 , 356–366.
Wright, S. D., Yong, C. G., Dawson, J. W., Whittaker, D. J.,
and Gardner, R. C. (2000). Riding the ice age El Niño?
Pacific biogeography and evolution of Metrosideros
subg.Metrosideros(Myrtaceae) inferred from nuclear
ribosomal DNA. Proceedings of the National Academy of
Sciences of the USA, 97 , 4118–4123.
Wright, S. J. (1980). Density compensation in island
avifaunas.Oecologia, 45 , 385–389.
Wyatt-Smith, J. (1953). The vegetation of Jarak Island,
Straits of Malacca. Journal of Ecology, 41 , 207–225.
Wylie, J. L. and Currie, D. J. (1993a). Species–energy the-
ory and patterns of species richness: I. Patterns of bird,
angiosperm, and mammal richness on islands. Biological
Conservation, 63 , 137–144.
Young, T. P. (1994). Natural die-offs of large mammals:
implications for conservation. Conservation Biology, 8 ,
410–418.
Zann, R. A. and Darjono (1992). The birds of Anak Krakatau:
the assembly of an avian community. Geojournal, 28 ,
261–270.
Zavaleta, E. S., Hobbs, R. J., and Money, H. A. (2001).
Viewing invasive species removal in a whole-ecosystem
context.Trends in Ecology and Evolution, 16 , 454–459.
Zavodna, M., Arens, P., van Dijk, P. J., Partomihardjo, T.,
Vosman, B., and van Damme, J. M. M. (2005).
Pollinating fig wasps: genetic consequences of island
recolonization. Journal of Evolutionary Biology, 18 ,
1234–1243.
Zimmerman, B. L. and Bierregaard, R. O., Jr (1986).
Relevance of the equilibrium theory of island biogeog-
raphy and species–area relations to conservation with a
case from Amazonia. Journal of Biogeography, 13 , 133–143.
Zink, R. M. (2002). A new perspective on the evolutionary
history of Darwin’s finches. Auk, 119 , 864–871.REFERENCES 381