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EARLY JURASSIC BIVALVE BIODIVERSITY 129

Fig. 2. Total extinction rates of bivalve species per million years through Early Jurassic time, (a) Andean
basins, (b) NW Europe. Key for time units as in Figure 1.


of cosmopolitans and the global extinction of

cosmopolitans. Taxa are classified as cosmopoli-

tans if they occurred in western South America

and/or NW Europe and in at least one

more region such as southern Europe, North

America, Siberia etc. Origination rates are

expressed as the number of species originating

in a substage divided by the duration of the

substage.

Finally, I calculated the rate at which species

immigrated into NW Europe and the Andean

basins, respectively. Immigration rates are

defined as the number of immigrating species

divided by the duration of the substage. To be

classified as an immigrant that utilized the

Hispanic Corridor for dispersal a species had to

be: (1) present at opposite sides of the Corridor;

(2) simultaneously absent in the western

Pacific/eastern Tethys; and (3) confined to

relatively low palaeolatitudes. Fulfilment of all

three criteria renders alternative dispersal

routes, such as migration around the periphery

of Pangaea or long-range dispersal of larvae

across the palaeo-Pacific, unlikely (see also

Aberhan 2001). For each bivalve species of the

two regions the geographic distribution was

determined by a survey of the relevant litera-

ture. This was greatly facilitated by the Jurassic

bivalve catalogue housed in Wurzburg. It

consists of copies of most figured specimens of

Jurassic bivalves along with locality and age

information.
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