Table 3.6. Summary data for tree species in permanent forest plots at Monteverde (see Haber,
"Descriptions," p. 90).
Site
a. Estacion Biologica
b. Elfin Forest
c. Nadkarni plots
Plot 1
Plot 3
All plots
d. Penas Blancas
e. San Gerardo
g. Stellar
h. Los Llanos
Area
(ha)
0.1
2 1 1 4 1 1 1 1
Families
27
NA
39
NA
47
39
36
NA
27
Species
41
NA
76
41
114
104
115
NA
62
Individuals
62
NA
585
531
1429
406
489
NA
644
Basal
Area
(mVha)
66
NA
66.3
NA
62
37.6
55.2
NA
34.6
Elevation
(m)
1600
1550
1450
1450
1450
750
1150
650
1200
Life Zone
Lower montane
wet/rain
Lower montane
rain
Lower montane
wet
Lower montane
wet
Lower montane
wet
Tropical wet
(transition)
Premontane
rain
Premontane
wet
Premontane
moist
Principal
Investigator
N. Nadkarni
R. Lawton
N. Nadkarni
N. Nadkarni
N. Nadkarni
W. Haber
M. Kerry and
W. Haber
E. Arevalo
E. Arevalo
NA = information not currently available.
habitat destruction. The dominance of montane spe-
cies in the region's flora results in large part from the
great diversity of orchids and ferns (both epiphytic
and terrestrial) in the cloud forest. Epiphytes comprise
29% of the flora and are most conspicuous and abun-
dant in the cloud forest (Table 3.7). The lower spe-
cies richness of the Atlantic slope may also reflect less
intensive collecting there.
Herbs. Herbs are defined here as erect, nonwoody
plants to distinguish them from vines (nonerect) and
2000
Figure 3.5. Number of species of ferns, monocots, and
dicots in the three general vegetation types (Pacific,
montane, and Atlantic) of the Monteverde area.
shrubs (woody). Some plants are intermediate among
these three categories; many pasture weeds (e.g., Agera-
tina and Sida.) develop woody stem bases, and some
ferns may grow either as epiphytes or terrestrial herbs.
These species were tabulated as their most common
growth form. A total of 616 species of terrestrial herbs
(including 144 ferns) (21% of the total flora) have been
identified in Monteverde (Table 3.7). This includes 245
species of dicotyledons in 44 families (13% of all dicot
species) and 227 monocotyledons in 21 families (28%
of all monocot species; Appendix 2). Herbaceous
species of dicots are most abundant in the families
Acanthaceae, Asteraceae, Gesneriaceae, Lamiaceae,
and Rubiaceae. The most diverse monocot families are
Araceae, Commelinaceae, Cyperaceae, Marantaceae,
Orchidaceae, and Poaceae (Fig. 3.6).
Many of the herb species, especially weedy mem-
bers of the Asteraceae and Poaceae, are more typical
of disturbed areas such as roadsides and pastures than
natural habitats. Some typical herbs of forest habitats
include species of Araceae (Dieffenbachia, Spathi-
phyllum), Cyclanthaceae (Carludovica, Cyclanthus),
Commelinaceae (Dichorisandra hexandra, Trades-
cantia zanonia), Costaceae (Costus), Gesneriaceae
(Alloplectus, Besleriaprinceps), Orchidaceae (Crani-
chis, Corymborkis), and Zingiberaceae (Renealmia).
Compared with wet forests at lower elevations, terres-
trial herbs form a smaller percentage of the biomass
in undisturbed montane habitats, occurring primarily
51 Plants and Vegetation