106 Invasive Stink Bugs and Related Species (Pentatomoidea)
these genera essentially possess those characters that define the tribe Cappaeini (see Section 2.2.10.7.11),
and it is fairly well accepted that Halyomorpha is closely related to Cappaea, the type genus of the tribe
Cappaeini. So, the genera listed above have been discussed under that tribe.
2.2.10.7.45.7 Banya Group This is a monotypic group containing the single genus and species
Banya leplaei Schouteden (Linnavuori 1982). Schouteden (1916) originally noted that this genus had
some resemblance to the genera Benia and Mabusana of the Veterna group (Cappaeini) (see Section
2.2.10.7.11) but differed strikingly by the sharp spine at the base of the abdomen, which extends onto the
metasternum. Members of the Cappaeini lack an abdominal spine, but several other groups, including
the Menidini (see Section 2.2.10.7.28), have a similar spine. In fact, Schouteden (1916) also suggested a
close relationship with the Menidini. Linnavuori (1982), however, gave a number of characters in which
Banya differs from members of the Menidini, but many of these may not be true if non-African meni-
dines are compared. As mentioned under that tribe, the Menidini is in dire need of revision and will
probably undergo many changes in the future.
2.2.10.7.45.8 Eipeliella Group Linnavuori (1982) proposed this group for a single genus, Eipeliella
Schumacher, which contains five African species. This genus originally was described as a member of the
Graphosomatinae (= Podopinae) (see Section 2.2.10.9), based primarily on the large scutellum (Schumacher
1912). We now know that an enlarged scutellum has evolved a number of times in the Pentatomoidea, and
that, by itself, is not enough to indicate relationship. Linnavuori (1982) indicated that the general body form
and the simple spermathecal bulb resembled that found in some members of the Eysarcorini (see Section
2.2.10.7.22); he then provided a number of characters that separate Eipeliella from eysarcorines. He finally
concluded that this genus might be related to the Antestiini (see Section 2.2.10.7.8).
2.2.10.7.45.9 Kelea Group This is another monotypic group containing a single genus and species,
Keleacoris congolensis (Distant). Kelea Schouteden is preoccupied, so Rider and Rolston (1995) proposed
Keleacoris as a replacement name. Linnavuori (1982) indicated that this species was similar in appearance to
species of Eurysaspis. The head is short and broad with the lateral margins of the juga distinctly notched ante-
rior to the compound eyes; the antenniferous tubercles are visible from dorsal view; and the apex of the head
is broadly truncate. The bucculae are rather broad. The rostrum extends to the mesocoxae. The pronotum is
convex, and the anterolateral margins are broadly convex and narrowly reflexed. The scutellum is somewhat
enlarged, ligulate, with distinct basal callosities; the frena are rather short. The base of the abdomen is armed
with a tumescence that meets with the posterior margin of the elevated metasternum. The mesosternum has
a medial carina only anteriorly. The female spermathecal bulb is simple without diverticula.
Linnavuori (1982) discussed possible connections with the Eurysaspini (see Section 2.2.10.7.21) but
ultimately concluded that this genus is probably a derivative of the Menida group (= Menidini) (see
Section 2.2.10.7.28), especially the genus Gwea Schouteden.
2.2.10.7.46 Other Unplaced or Questionably Placed Genera
Gross (1975b, 1976) and Linnavuori (1982) dealt with nearly all of the difficult Australian and African
genera, placing them into various generic groups. There are, however, a number of pentatomid genera
from the Orient and Neotropics that have never been placed in any known tribes, and their current place-
ment remains unknown. Further study is needed before proper placement can be made confidently. We
provide a few comments that may help with future placement.
Araducta Walker, 1867
Walker (1867b) originally described this genus as having a pectoral keel that extended to the fore coxae,
and he compared this genus with Cuspicona (Figure 2.23L), a member of the Rhynchocorini. Distant
(1900b) redescribed the genus, acknowledging the pectoral keel, but made some changes in describing
the abdominal spine; he placed this genus near the genus Brachystethus, a member of the Edessinae.
Distant also described or transferred four more species in Araducta, all of which have now been trans-
ferred to either Dabessus or Dunnius, genera now tentatively placed in the Menidini. Kirkaldy (1909)
catalogued this genus after the genus Exithemus and before the genus Lelia, members of the tribes