Higher Systematics of the Pentatomoidea 107
Rolstoniellini and Pentatomini, respectively. We have examined specimens and confirm that they pos-
sess the sternal structure that is characteristic of the rhynchocorines; although the coloration of these
specimens differs from the typical, we still treat this genus as a member of the Rhynchocorini (see
Section 2.2.10.7.38). The genus occurs in the Malay Archipelago and New Guinea.
Banasa Stål, 1860; Disderia Bergroth, 1910; Elanela Rolston, 1980; Glaucioides Thomas, 1980;
Grazia Rolston, 1981; Janeirona Distant, 1911; Kermana Rolston, 1981; Modicia Stål, 1872;
Nocheta Rolston, 1980; Pallantia Stål, 1862; Rio Kirkaldy, 1909; and Vidada Rolston, 1980
For New World genera, Rolston et al. (1980) divided his broadly conceived Pentatomini into three sec-
tions. Section one contained those genera in which the abdominal venter was unarmed basally; section
two contained those genera in which the base of the abdomen was armed with a spine or tubercle that
was unapposed by an elevated metasternum (i.e., the apex was free distally); and section three contained
those genera in which the abdominal venter was armed with a tumescence, tubercle, or small spine that
was apposed by an elevated metasternum and might be truncate or notched.
Nearly all of the Old World genera that have the abdominal venter armed would fall into Rolston’s
section two (Rolston and McDonald 1981); that is, the abdominal armature is unapposed by the meta-
sternum. There are however, a number of New World genera that belong in Rolston’s section three
(Rolston et al. 1980). Some of these have been moved elsewhere (e.g., Arvelius into the Chlorocorini;
Brachystethus and Lopadusa into the Edessinae), or at least have been discussed elsewhere in this chapter
(e.g., Evoplitus and related genera). Many of the remaining genera do not fit neatly into the well-known
tribes that have the abdomen armed (e.g., Catacanthini, Eurysaspini, Menidini, Nezarini, Piezodorini).
Based on a similar general appearance, we treat Elanela (Figure 2.22C) and Rio (Figure 2.22E) as
members of the Menidini (maybe erroneously). We believe the treatment of the remaining genera as
members of the Pentatomini is suspicious, but, without further evidence, we conservatively leave them in
the Pentatomini. Several of the genera treated here have been revised recently: Banasa (Figure 2.31D)
(Thomas and Yonke 1981, 1988, 1990), Disderia (Rolston 1983b), Elanela (Grazia and Greve 2011,
Grazia et al. 2016a), Pallantia (Figure 2.23E) (Grazia 1980a, 1986; Brailovsky 1986), and Rio (Grazia
and Fortes 1995, Fortes and Grazia 2000). The monotypic genus Glaucioides Thomas (Figure 2.23A)
was described recently (Rolston et al. 1980).
Brachyhalys Ahmad, 1981
Ahmad (1981) listed as a new genus and species Brachyhalys pilosum, and he provided a new family-level
name, Brachyhalyini, for its inclusion. He did not, however, provide any descriptions of the tribe, genus, or
species, and there were no type designations. As such, these names should all be considered as unavailable.
Brasilania Jensen-Haarup, 1931
We have not examined any specimens of this genus. Jensen-Haarup (1931a) originally placed this
New World genus near the Old World genus Bathycoelia (currently in the Bathycoeliini or possibly
Pentatomini; see Section 2.2.10.7.10). Jensen-Haarup (1931a) described this genus as having the abdom-
inal venter unarmed, and Rolston (1987a) included it in an artificial group of seven South American gen-
era of Pentatomini that have an unarmed abdominal venter and elongate ostiolar rugae. The sternum was
originally described as not carinate. This genus is unusual in that the coloration is described as pitchy
black all over (including the membranes), and the rostrum is extremely long, reaching well beyond the
apex of the abdomen.
Cachaniellus Rider, 2007
This genus originally was described by Cachan (1952) using the preoccupied generic name Sambirania.
He placed it in a broadly conceived Carpocorini but indicated that it was related to Corisseura, the place-
ment of which also is uncertain. We tentatively treat both genera as members of the Eysarcorini. Both
genera are endemic to Madagascar.
Catalampusa Breddin, 1903
Breddin (1903) described its only known species, C. oenops Breddin, from Bolivia, placing it between a
new species of Nezara and a new species of Tibilis. In the original description, Breddin (1903) indicated