108 Invasive Stink Bugs and Related Species (Pentatomoidea)
that the metasternum is elevated and received the short abdominal spine, thus placing it in Rolston’s
section three. Kirkaldy (1909), the last worker to mention this genus, also catalogued it near the genus
Tibilis, thus perhaps indicating a possible relationship with that genus. Because Tibilis is a member of the
Pentatomini, we are tentatively treating this genus as a member of this tribe.
Chrysodarecus Breddin, 1903
Breddin (1903) did not place this genus in any known tribe or subfamily, nor did he mention any possible
relationships. Kirkaldy (1909) catalogued it among the Discocephalinae, possibly because the next genus
dealt with in Breddin’s work was Adoxoplatys, a member of the Discocephalinae. In truth, the original
description in some ways is quite similar to the genus Placocoris Mayr. Breddin (1903) stated that the
body was quite flat and had a double row of spines on the hind femora (also found in Placocoris). The
single species of this genus was described from Peru.
Cyptocephala Berg, 1883; Te p a Rolston and McDonald, 1984; and Thyanta Stål, 1862
These three New World genera are obviously related to each other and although they have been studied
extensively by one of us (DAR), their tribal placement remains questionable. Rolston and McDonald
(1984), by virtue of their having an unarmed abdominal venter, placed these genera in section one of
their broadly conceived Pentatomini. These three genera also have the ostiolar rugae relatively elongate,
reaching to (Tep a) or extending beyond (Cyptocephala and Thyanta [Figure 2.32L]) the middle of the
metapleura, with the apices usually acuminate. Because the adults are usually green (yellowish or brown
forms are present in colder months), and usually remain green after death, we have tentatively placed
them in the Nezarini. Further study is needed to confirm this placement. All three genera have been
revised: Cyptocephala (Rolston (1986), Tep a (Rolston 1972, Rider 1986), and Thyanta (Figure 2.32L)
(Rider and Chapin 1991, 1992). As a side note, it appears that at least some of the subgenera of Thyanta
probably should be elevated to generic rank.
Dardjilingia Yang, 1936 and Zouicoris Zheng, 1986
These two genera were described from India and Bhutan, and from southwestern China, respectively;
both have remained monotypic so far. They share an unarmed abdominal venter; a mesosternum provided
with a very low median carina; a relatively short ostiolar ruga not reaching the middle of the metapleuron;
and a rostrum that is relatively short, approaching or reaching the middle coxae. The male genitalia and
the female genital plates of members of the two genera are similar as well, and there is little doubt that
they belong to the same tribe. It is difficult, however, to determine which tribe this should be because
the general habitus of these species is quite different from the majority of pentatomids. Yang (1936)
compared Dardjilingia with Neogynenica, currently in the Lestonocorini (see Section 2.2.10.7.25); the
female genital plates, however, are much different from the condition found in members of that tribe (also
see Salini 2016). Zheng (1986) indicated that the short rostrum in Zouicoris might ally this genus with
the phyllocephalines, but he ultimately stated that it was not a member of that group and that it probably
belonged in the Pentatominae; he did not, however, indicate to which tribe it might belong.
Lakhonia Yang, 1936
This genus, containing a single species occurring in Thailand and Southwest China, never has been for-
mally placed in any pentatomid subfamily or tribe. It is obvious from the illustrations and description that
the more recently described genus, Oedocoris Zheng and Liu, is synonymous (Rider et al. 2002), and yet its
tribal placement remains unknown. The dorsal surface is covered by a thick layer of short, soft, hairs, and
the head and anterior disk of the pronotum are strongly declivent. The abdominal venter is unarmed, and
the ostiolar rugae are short and auriculate. This suite of characters is similar to members of the Carpocorini.
Leovitius Distant, 1900
Distant (1900b) originally related this genus to the genus Prionochilus (a preoccupied name now placed
in the synonymy of the pentatomine genus Lelia). He later (1902) placed Leovitius in the Rhynchocorini,
even though Dallas (1851) and Distant, himself (1902), described the ventral armature as consisting of a
long basal abdominal spine, reaching beyond the front coxae. There is no mention of the elevated meta-
sternum (meeting posteriorly with the abdominal spine or tubercle), or the mesosternum being elevated