112 Invasive Stink Bugs and Related Species (Pentatomoidea)
prägen müssen” [Despite of it I have named 22 species, many times I had also to create new generic
names]. This could be interpreted as his intention to describe the new genera in combination, and may
allow for Halynoides, Pseudopalomena, Rhomboidea, and Suspectocoris to be valid. At any rate, he did
not indicate to what tribe these species belonged. Suspectocoris grandis Jordan originally was placed
in the Acanthosomatidae, but Popov (2007) transferred it to the Pentatomidae. Jordan (1967) originally
may have intended to show relationship by the names he chose (i.e., Carpocoroides might be related to
Carpocoris, Halynoides might be related to Halys, and so on), but we do not know this for sure. Popov
(2007) indicated that Pentatomoides might actually be a synonym of the extant genus Carpocoris, and
he transferred one of Jordan’s Pentatomoides species to the genus Carpocoroides.
Deryeuma Piton, 1940
Piton (1940) originally indicated that this genus was similar to the strachiine genus Eurydema (notice
that Deryeuma is an anagram of Eurydema).
Mesohalys Beier, 1952
Beier (1952) originally placed this genus, and its only included species, M. munzenbergiana Beier, in the
Halyaria (= Halyini).
Necanicarbula Zhang, 1989
This fossil genus originally was placed near the eysarcorine genera Carbula and Dymantiscus Hsiao
(Zhang 1989).
Poliocoris Kirkaldy, 1910 and Teleocoris Kirkaldy, 1910
Kirkaldy (1910) originally indicated that Teleocoris had some resemblance to various tessaratomids but
ultimately concluded that it probably belonged in the Pentatomidae: Halyini. He further indicated that
Poliocoris probably was related to Teleocoris and, thus, it should also belong in the Halyini.
Taubatecoris Martins-Neto, 1997
This is another fossil genus, recently described from Brazil (Martins-Neto 1997). It originally was not
placed in any known tribe; its correct taxonomic placement requires further study.
2.2.10.8 Phyllocephalinae Amyot and Serville, 1843
This interesting group seems to have received little attention in recent years. It is an Old World group,
occurring in Africa, the Middle East, the Indian subregion, the Oriental region, and extending into
Australia; it currently contains 45 genera and 214 species (Tables 2.2, 2.3).
This taxon was proposed first at the family-level (Phyllocéphalides) by Amyot and Serville (1843)
and was recognized as a family (Phyllocephalidae) by Dallas (1851). This group through time has been
treated at all levels from a genus group (Gross 1976), a tribe (Leston 1952a) within the Pentatominae, a
subfamily (Stål, 1865, Lethierry and Severin 1893, Distant 1902, Kirkaldy 1909, China and Miller 1959,
Linnavuori 1982, Schuh and Slater 1995, Rider 2006a), or a family (Vidal 1949, Miyamoto 1961). Gross
(1976) treated them as a genus group at the same level as other genus groups (= pentatomine tribes) and
related this group to the Cephaloplatus group (based on similar jugal expansions of the head, ostiolar
rugae, and even male genital characters). It appears that the general consensus among recent authors is
to treat it as a subfamily.
Members of this subfamily are medium to large in size, some are ovate (Figure 2.27G) whereas oth-
ers are elongate, almost parallel-sided (Figure 2.27H), and the dorsal surface is often quite rugulose;
their coloration is usually yellows, tans, or browns, allowing them to blend in with their surroundings.
The juga are usually longer than the tylus, sometimes meeting in front of the tylus, and may be expanded
or foliate; there sometimes is a tooth-like projection anterior to each compound eye. The bucculae are
usually short, flap-like (Figure 2.1O). The single most diagnostic character is the distinctively short
rostrum, which does not or only barely surpasses the procoxae (it may be a little longer in Australian
genera, but it still reaches, at most, to the anterior margin of the mesocoxae) (Figure 2.5B). Rostral seg-
ment I and half of segment II lie between the bucculae. The humeral angles may be rounded or greatly