25

2

Higher Systematics of the Pentatomoidea

David A. Rider, Cristiano F. Schwertner, Jitka Vilímová,
Dávid Rédei, Petr Kment, and Donald B. Thomas

CONTENTS

2.1 Introduction .................................................................................................................................... 27
2.2 Pentatomoidea ................................................................................................................................ 29
2.2.1 Key to the Families of Pentatomoidea............................................................................... 29
2.2.2 Acanthosomatidae Signoret, 1863 ..................................................................................... 34
2.2.2.1 Key to the Subfamilies of Acanthosomatidae .................................................. 38
2.2.3 Canopidae Amyot and Serville, 1843 ................................................................................ 38
2.2.4 Cydnidae Billberg, 1820 .................................................................................................... 39
2.2.4.1 Key to the Subfamilies of Cydnidae .................................................................. 41
2.2.5 Dinidoridae Stål, 1868 ....................................................................................................... 42
2.2.5.1 Key to the Subfamilies of Dinidoridae ............................................................. 44
2.2.6 Lestoniidae China, 1955 .................................................................................................... 44
2.2.7 Megarididae McAtee and Malloch, 1928 .......................................................................... 46
2.2.8 Mesopentacoridae Popov, 1968 ......................................................................................... 47
2.2.9 Parastrachiidae Oshanin, 1922 .......................................................................................... 47
2.2.10 Pentatomidae Leach, 1815 ................................................................................................. 48
2.2.10.1 Key to the Subfamilies of Pentatomidae ........................................................... 51
2.2.10.2 Aphylinae Bergroth, 1906 ................................................................................ 54
2.2.10.3 Asopinae Amyot and Serville, 1843 ................................................................. 54
2.2.10.4 Cyrtocorinae Distant, 1880 .............................................................................. 57
2.2.10.5 Discocephalinae Fieber, 1860 ........................................................................... 57
2.2.10.5.1 Key to the Tribes of Discocephalinae .......................................... 59
2.2.10.6 Edessinae Amyot and Serville, 1843 ................................................................ 59
2.2.10.7 Pentatominae Leach, 1815 ................................................................................ 60
2.2.10.7.1 Tentative Key to the Tribes of Pentatominae ................................ 61
2.2.10.7.2 Aeliini Douglas and Scott, 1865 ................................................... 69
2.2.10.7.3 Aeptini Stål, 1871 ......................................................................... 69
2.2.10.7.4 Aeschrocorini Distant, 1902 ........................................................ 70
2.2.10.7.5 Agaeini Cachan, 1952 ................................................................... 71
2.2.10.7.6 Agonoscelidini Atkinson, 1888 .................................................... 72
2.2.10.7.7 Amyntorini Distant, 1902 ............................................................ 72
2.2.10.7.8 Antestiini Distant, 1902 ............................................................... 73
2.2.10.7.9 Axiagastini Atkinson, 1888 .......................................................... 74
2.2.10.7.10 Bathycoeliini Atkinson, 1888 ....................................................... 75
2.2.10.7.11 Cappaeini Atkinson, 1888 ............................................................ 76
2.2.10.7.12 Carpocorini Mulsant and Rey, 1866............................................. 76
2.2.10.7.13 Catacanthini Atkinson, 1888 ........................................................ 79
2.2.10.7.14 Caystrini Ahmad and Afzal, 1979................................................ 80
2.2.10.7.15 Chlorocorini Rider, Greve, Schwertner, and Grazia, New Tribe .... 80