Invasive Stink Bugs and Related Species (Pentatomoidea)

Higher Systematics of the Pentatomoidea 45

(1991), and Schaefer (1993) seemed to support a relationship with the Plataspidae, but they ultimately
decided that there were enough differences to maintain separate family status. The phylogenetic studies
by Grazia et al. (2008) and Wu et al. (2016) provide further support for the recognition of Lestoniidae as
the sister group of the Acanthosomatidae.
Members of this family are relatively small (5-6 mm), ovoid, dorsally convex, and ventrally flattened
(Schuh and Slater 1995). The scutellum is large in Lestonia haustorifera (Figure 2.25D) covering much,

ABC

DE

F G

FIGURE 2.4 A, Coridius viduatus, evaporatorium, ventral view; B, Coridius viduatus, ostiolar ruga, ventral view;
C, Tessaratoma papillosa, ostiolar ruga, ventral view; D, Oncomeris flavicornis, ostiolar ruga, ventral view; E, Eumenotes
obscura, thorax and anterior abdomen, lateral view; F, Kayesia parva, evaporatorium, ventral view; G, Eurygaster maura,
evaporatorium, ventral view.