Herrera, E. A. 1986. The behavioural ecology of the capybara
Hydrochoerus hydrochaeris. PhD diss., University of Oxford.
———. 1992. Size of testes and scent glands in capybaras, Hy-
drochaeris hydrochaeris(Rodentia, Caviomorpha). Journal of
Mammalogy73 : 871–75.
———. 1998. Reproductive strategies of female capybaras: Dry
season gestations. Symposium of the Zoological Society of
London71 : 281– 92.
Herrera, E. A., and D. W. Macdonald. 1987. Group stability
and the structure of a capybara population. Symposium of the
Zoological Society of London58 : 1–350.
———. 1989. Resource utilization and territoriality in group-
living Capybaras Hydrochoerus hydrochaeris. Journal of
Animal Ecology58 : 667–79.
———. 1993. Aggression, dominance and mating success among
capybara males (Hydrochoerus hydrochaeris). Behavioural
Ecology4 : 114 –19.
———. 1994. Social significance of scent marking in capybaras.
Journal of Mammalogy75 : 410 –15.
Herrmann, B., M. Bucan, P. E. Mains, A. M. Frischauf, L. M.
Silver, and H. Lehrach. 1986. Genetic analysis of the proxi-
mal portion of the mouse tcomplex: Evidence for a second
inversion within thaplotypes. Cell44 : 469 –76.
Herrmann, B. G., B. Koschorz, K. Wertz, K. J. McLaughlin, and
A. Kispert. 1999. A protein kinase encoded by the tcomplex
responder gene causes non-mendelian inheritance. Nature
402 : 141– 46.
Herron, M. D., T. A. Castoe, and C. L. Parkinson. 2004. Sciurid
phylogeny and the paraphyly of Holarctic ground squirrels
(Spermophilus). Molecular Phylogenetics and Evolution
31 : 1015 –30.
Hersek, M. J. 1990. Behavior of predator and prey in a highly
coevolved system: Northern Pacific rattlesnakes and Califor-
nia ground squirrels. PhD diss., University of California,
Davis.
Hersek, M. J., and D. H. Owings. 1993. Tail flagging by adult
California ground squirrels: A tonic signal that serves differ-
ent functions for males and females. Animal Behaviour
46 : 129 –38.
———. 1994. Tail flagging by young California ground squirrels,
Spermophilus beecheyi: Age-specific participation in a tonic
communicative system. Animal Behaviour48 : 803 –11.
Hershkovitz, P. 1962. Evolution of neotropical cricetine rodents
(Muridae) with special reference to the Phyllotine group. Fiel-
diana Zoology46 : 1–524.
———. 1966. Mice, land bridges and Latin American faunal in-
terchange. In Ectoparasites of Panama,ed. R. L. Wenzel and
V. J. Tipton, 725 –51. Chicago: Field Museum of Natural
History.
Herzog, H. A., and G. M. Burghardt. 1986. Development of
antipredator responses in snakes: I. Defensive and open-field
behaviors in newborns and adults of three species of garter
snakes (Thamnophis melanogaster, T. sirtalis, T. butleri).
Journal of Comparative Psychology100 : 372 –79.
Heske, E. J. 1987. Pregnancy interruption by strange males in the
California vole. Journal of Mammalogy68 : 406 –10.
Heske, E. J., and R. S. Ostfeld. 1990. Sexual dimorphism in size,
relative size of testes, and mating systems in North American
voles. Journal of Mammalogy71 : 510 –19.
Heske, E. J., R. S. Ostfeld, and W. Z. Lidicker, Jr. 1984. Compet-
itive interactions between Microtus californicusand Rei-
throdontomys megalotisduring two peaks of Microtusabun-
dance. Journal of Mammalogy 65 : 271– 80.
Hess, G. R. 1994. Conservation corridors and contagious dis-
ease: A cautionary note. Conservation Biology8 : 256 – 62.
Hestbeck, J. B. 1982. Population regulation of cyclic mammals:
The social fence hypothesis. Oikos39 : 157– 63.
Heth, G. 1989. Burrow patterns of the mole rat Spalax ehren-
bergiin two soil types (terra rossa and rendzina) in Mount
Carmel. Israel Journal of Zoology7 : 39 –56.
———. 1991. The environmental impact of subterranean mole
rats (Spalax ehrenbergi) and their burrows. Symposia of the
Zoological Society, London63 : 265 – 80.
Heth, G., G. Beauchamp, E. Nevo, and K. Yamazaki. 1996. Spe-
cies, population and individual specific odors in urine of mole
rats (Spalax ehrenbergi) detected by laboratory rats. Chemoe-
cology7 : 107–11.
Heth, G., A. Beiles, and E. Nevo. 1988. Adaptive variation of
pelage color within and between species of the subterranean
mole rat (Spalax ehrenbergi)in Israel. Oecologia74: 617–22.
Heth, G., E. Frankenberg, and E. Nevo. 1986. Adaptive optimal
sound for vocal communication in tunnels of a subterranean
mammal (Spalax ehrenbergi). Experientia42 : 1287– 89.
———. 1988. “Courtship” call of subterranean mole rats
(Spalax ehrenbergi): Physical analysis. Journal of Mammalogy
69 : 121–25.
Heth, G., E. Frankenberg, H. Pratt, and E. Nevo. 1991. Seismic
communication in the blind subterranean mole rats: Patterns
of head thumping and of their detection in the Spalax ehren-
bergisuperspecies in Israel. Journal of Zoology, London
224 : 633 –38.
Heth, G., E. Frankenberg, A. Raz, and E. Nevo. 1987. Vibrational
communication in subterranean mole rats (Spalax ehrenbergi).
Behaviorial Ecology and Sociobiology21 : 31–33.
Heth, G., E. Nevo, R. Ikan, V. Weinstein, U. Ravid, and H. Dun-
can. 1992. Differential olfactory perception of enantiomeric
compounds by blind subterranean mole rats (Spalax ehren-
bergi). Experientia48 : 897– 902.
Heth, G., and J. Todrank. 1995. Assessing chemosensory percep-
tion in subterranean mole rats: Different responses to smelling
versus touching odorous stimuli. Animal Behaviour49 :
1009 –15.
———. 2000. Individual odour similarities across species paral-
lel phylogenetic relationships in the S. ehrenbergisuperspecies
of mole-rats. Animal Behaviour2000 : 789 – 95.
Heth, G., J. Todrank, S. Begall, R. Koch, Y. Zilbiger, E. Nevo,
S. H. Braude, and H. Burda. 2002. Odours underground:
Subterranean rodents may not forgage “blindly.” Behavioral
Ecology and Sociobiology52 : 53 –58.
Heth, G., J. Todrank, and H. Burda. 2002. Individual odor simi-
larities within colonies and across species of Cryptomysmole
rats. Journal of Mammalogy83 : 569 –75.
Heyes, C. M. 1993. Imitation, culture and cognition. Animal
Behaviour46 : 999 –1010.
———. 1996. Genuine imitation? In Social learning and imita-
tion: The roots of culture,ed. C.M. Heyes and B.G. Galef,
Jr., 371– 90. New York: Academic Press.
Heyes, C. M., and B. G. Galef, Jr. 1996.Social learning in
animals: The roots of culture.San Diego, CA: Academic
Press.
Heyes, C. M., and G. R. Dawson. 1990. A demonstration of
observational learning using a bidirectional control. Quar-
terly Journal of Experimental PsychologyB 42 : 59 –71.
Heyes, C. M., G. R. Dawson, and T. Nokes. 1992. Imitation in
rats: Initial responding and transfer evidence. Quarterly Jour-
nal of Experimental PsychologyB 45 : 229 – 40.
532 References