316 Ordinary Differential Equations
system is a simple example of a linear system with variable coefficients (a
linear system in which not all of the coefficient functions are constant).
(6)
y; = - 3y1 + (x + l/x)y2 + sinx.
Here a 11 (x) = x^2 , a 12 (x) =-ex, b 1 (x) = 25, a21(x) = -3, a22(x) = x + 1/x,
and b2(x) = sinx.
DEFINITIONS Homogeneous, Nonhomogeneous, and Nonlinear
Systems of First-Order Equations
If all of the functions bi ( x) are identically zero, then the linear system ( 5)
is said to be homogeneous.
If any bi(x) is not identically equal to zero, then the linear system (5) is
said to be nonhomogeneous.
All systems of first-order differential equations not having the form of
system (5) are called nonlinear systems.
As an example of a nonlinear system of differential equations, we introduce
the Volterra prey-predator model, one of the fundamental models of mathe-
matical ecology. Let y 1 ( x) represent the prey population at time x and let
Y2 ( x) represent the predator population at time x. In the absence of predators,
the population y 1 is assumed to grow according to the Malthusian population
model: dyif dx = ay 1 where a > 0 is the growth rate. The loss of population
due to predation is assumed to be proportional to the number of encounters
between prey and predator- that is , proportional to the product y 1 y 2. Thus,
the rate of change of the prey population becomes dyif dx = ay1 -by 1 Y2 where
b > 0 is a constant which represents the proportion of encounters between the
prey and predators which result in death to the prey. Without the prey to feed
upon, it is assumed that the predators would die off according to the Malthu-
sian population model: dy2/ dx = -cy2 where c > 0 is the death rate. Due
to predation, the predator population is assumed to grow at a rate which is
proportional to the number of encounters between prey and predator. Hence,
the rate of change in predator population becomes dy 2 /dx = - cy 2 + dy 1 y 2
where d > 0 is a constant which represents the proportion of encounters be-
tween prey and predator which is beneficial to the predator. Therefore, the
Volterra prey-predator model is