140 CHAPTER 6 | Macroevolution and the Early Primates
Miocene Apes and
Human Origins
As described above, determining which Miocene apes were
directly ancestral to humans is one of the key questions in
primate evolution. Molecular evidence directs our attention
to Africa between 5 and 8 mya (Figure 6.8). Though any
fossil discoveries in Africa from this critical time period
that it indicates that the gorilla lineage had become dis-
tinct 2 to 4 million years earlier than that. The species is
called Chororapithecus abyssinicus, after Chorora, the local
area where the fossil was found, and Abyssinia, the ancient
name of Ethiopia. While Ethiopia has been the home for
numerous spectacular fossil specimens that are part of the
human lineage, this is the first fossil ape discovery from
this region. Other scientists do not concur, however, and
feel more fossil evidence is warranted before pushing back
the timing of the split.
Millions of years ago
0
5
10
15
20
25
30
35
40
45
OLIGOCENE
MIOCENE
Aegyptopithecus
Proconsul
Sivapithecus
Baboons Macaques Cercopithecus Presbytis Colobus Gibbon-
Siamang
Orangutan Gorilla ChimpanzeeHomo
CERCOPITHECOIDS HOMINOIDS
CATARRHINI
Orrorin
tugenensis?
Sahelanthropus
tchadensis?
(Toumai)
Chororapithecus
abyssinicus?
Australo-
pithecus
Figure 6.8 Although debate continues over details, this chart represents a reasonable
reconstruction of evolutionary relationships among the Old World anthropoid primates. (Extinct
evolutionary lines are not shown.) The 2007 discovery of a fossil ancestor to gorillas has suggested a
new interpretation of the timing and nature of the split between humans and the African apes.
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