CHAPTER 31
The Heart as a Pump 509
EARLY DIASTOLE
Once the ventricular muscle is fully contracted, the already
falling ventricular pressures drop more rapidly. This is the pe-
riod of
protodiastole,
which lasts about 0.04 s. It ends when
the momentum of the ejected blood is overcome and the aortic
and pulmonary valves close, setting up transient vibrations in
the blood and blood vessel walls. After the valves are closed,
pressure continues to drop rapidly during the period of
iso-
volumetric ventricular relaxation.
Isovolumetric relaxation
ends when the ventricular pressure falls below the atrial pres-
sure and the AV valves open, permitting the ventricles to fill.
Filling is rapid at first, then slows as the next cardiac contrac-
tion approaches. Atrial pressure continues to rise after the end
of ventricular systole until the AV valves open, then drops and
slowly rises again until the next atrial systole.
FIGURE 31–1
Divisions of the cardiac cycle: A) systole and B) diastole.
The phases of the cycle are identical in both halves of the heart.
The direction in which the pressure difference favors flow is denoted by an arrow; note, however, that flow will not actually occur if valve prevents it.
SystoleA
B
Blood flows out of ventricle
Ventricles
contract
Atria
relaxed
Ventricles
relaxed
Atria
relaxed
Ventricles
relaxed
Atria
relaxed
Ventricles
relaxed
Atria
contract
Ventricles
contract
Atria
relaxed
Closed
Closed Closed
Open
Diastole
Blood flows into ventricles
Atrial contraction
AV valves:
Aortic and
pulmonary valves:
Closed
Closed Open
Closed
Open
Closed
AV valves:
Aortic and
pulmonary valves:
Isovolumetric ventricular contraction
Isovolumetric ventricular relaxation
Ventricular ejection
Ventricular filling