HUMAN BIOLOGY

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the RespiRatoRy system 183

expiration Exhalation of
a breath.
inspiration Inhalation of
a breath.
respiratory cycle The in/
out pattern of breathing.
tidal volume Volume of
air entering and leaving
the lungs during a normal
breath.
vital capacity Maximum
volume of air that can move
out of the lungs after the
deepest possible inhalation.

Pneumothorax, or a “collapsed lung,” is caused by
injury or illness that allows air to enter the pleural cavity.
The lungs can’t expand normally and breathing becomes
difficult and painful.

how much air is in a “breath”?
About 500 milliliters (two cupfuls) of air enters or leaves
your lungs in a normal breath. This volume of air is
called tidal volume. You can increase the amount of air
you inhale or exhale, however. In addition to air taken in
as part of the tidal volume, a person can forcibly inhale
roughly 3,100 milliliters of air, called the inspiratory reserve
volume. By forcibly exhaling, you can expel an additional
expiratory reserve volume of about 1,200 milliliters of air.
Vital capacity is the maximum vol-
ume of air that can move out of the
lungs after you inhale as deeply
as possible. It is about 4,800 milli-
liters for a healthy young man and
about 3,800 milliliters for a healthy
young woman. As a practical matter,
people rarely take in more than half
their vital capacity, even when they
breathe deeply during strenuous
exercise. At the end of your deepest
exhalation, your lungs still are not
completely emptied of air; another
roughly 1,200 milliliters of residual
volume remains (Figure 10.10).
How much of the 500 milliliters of inspired air is
available for gas exchange? Between breaths, about
150 milliliters of exhaled “dead” air remains in the air-
ways. Thus only about 350 (500 2 150) milliliters of fresh
air actually reaches the alveoli each time you inhale. An
adult typically breathes at least twelve times per minute.
This rate of ventilation supplies the alveoli with 4,200
(350 3 12) milliliters of fresh air every 60 seconds—about
the volume of soda pop in four 1-liter bottles.
When food “goes down the wrong way” and enters the
trachea (instead of the esophagus), it’s impossible to inhale
or exhale normally. A choking person can suffocate in
just a few minutes. The Heimlich maneuver can dislodge
material from the trachea by elevating the diaphragm
(Figure 10.11). This reduces the chest volume, forcing air up
the trachea. With luck, the air will rush out with enough
force to eject the item.

0

1,000

2,000

3,000

4,000

5,000

Lung volume (milliliters)

6,000

Time

inspiratory
reserve
volume

expiratory
reserve volume

tidal
volume

vital
capacity

total
lung
capacity

residual
volume

Figure 10.10 Animated! Lung volume changes during quiet
breathing and during forced inspiration and expiration. In this
graph you can see “spikes” above and below the normal tidal
volume. (© Cengage Learning)


Figure 10.11 Animated! The Heimlich maneuver is an
emergency procedure designed to rescue a choking adult.
(Top: © Cengage Learning 2015; Bottom: From Starr/ Taggart/Evers/Starr, Biology,
12E. © 2009 Cengage Learning)
What is the respiratory cycle?



  • The in/out air movements of breathing make up the
    respiratory cycle.

  • Breathing air movements occur when lung volume changes as
    the chest cavity expands and shrinks.

  • These changes alter the pressure gradients between the lungs
    and outside air.


takE-homE mEssaGE

You may recall from Chapter 2 that the hydrogen bonds
between water molecules prevent them from being easily
pulled apart. This cohesiveness of water molecules in the
fluid in the pleural sac also helps your lungs hug the chest
wall, in much the same way that two wet panes of glass
resist being pulled apart.


a Back blows. Have the person lean forward. Use the heel of
your hand to strike between the shoulder blades.

B Abdominal thrusts. Stand behind the person and place one
fist below the rib cage, just above the navel, with your thumb
facing inward. Cover the fist with your other hand and thrust
inward and upward with both fists.

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