850 THE STRUCTURE OF EVOLUTIONARY THEORY
also strongly implicating punctuated equilibrium as the major generator of larger
trends, if only because the fruits of anagenesis get plucked so quickly unless they
can be "locked up" in cladogenetic iterations.
The "dissection" of punctuations to infer both existence
and modality
Once a literal punctuation has been noted, and a cladogenetic origin inferred by
such criteria as the documentation of ancestral survival, further testing of
punctuated equilibrium as the mode of origin for the new species may be achieved
by several standards that might be characterized (somewhat metaphorically)
9 - 22. In other areas of the range, as here in South Dakota, only the descendant species lives
during the entire interval.