498 MHR • Unit 5 Population Dynamics
a particular habitat. For example, they might be
specialists at finding, hunting, or eating a particular
type of food, or at physiologically coping with a
particular temperature and moisture regime.
In a stable environment, individuals that
produce a few large offspring that are cared for to
ensure their survival are favoured. They are more
likely to pass their genes along to subsequent
generations. An organism like this is said to have
an equilibrial life history. This type of life history
is also called K-selected, since it is seen in
populations that are near Kin size. In these
environments, K-selectionis at work.
As always, it is important to realize that the
distinction between these two general types of life
history is not absolute. Real environments are
highly complex. So, too, are the selective forces
that act within those environments. Most
organisms therefore, experience both r- and
K-selection to a greater or lesser extent, and many
display features of both types of life histories. It is
probably useful to think of purely opportunistic
(r-selected) and purely equilibrial (K-selected) life
histories as forming opposite ends of a theoretical
continuum. Most populations lie somewhere
between these two extremes. In the Thinking Lab
on page 497, you will illustrate your understanding
of the differences between an opportunistic life
history and an equilibrial life history.
Looking Ahead
Human life history features are closer to what
ecologists would consider the equilibrial or
K-selected end of the scale. This suggests that
for most of its evolutionary history, the human
population has been at or near its carrying capacity.
Several lines of evidence from archeologists,
anthropologists, and historians support this
viewpoint.
However, as will be discussed in the next
chapter, about 8000 years ago humans managed to
change their environment’s carrying capacity. The
human population then entered into a period of
rapid growth that is still underway. Much of the
biosphere has been affected by this growth, and the
ecology of many species has been altered as a
result. The future survival of an array of species,
including Homo sapiens, may depend on whether
the growth of the human population and its impact
on Earth can be controlled.
SECTION REVIEW
- Identify each of the elements in the (K-N)/N
portion of the logistic growth equation (that is, what
does each of the letters represent?). How does this
portion of the equation affect the overall rate of
growth of a population? - Distinguish between opportunistic and
equilibrial life histories. - Distinguish between r-and K-selection.
- What factors typically limit the population size
of species with very high birth rates? - Draw a survivorship curve for a population in which
most of the mortality occurs among the youngest
members of the species. Provide examples of two
species that demonstrate such a survivorship pattern. - What stage of an organism’s life cycle typically
has the highest fecundity rate? Explain your answer. - Do humans typically have only one reproductive
event or several over their lifetimes? Explain your
answer. - What is the survivorship pattern for Canada’s
human population?
9. Compare the typical age structures seen in
populations with Type I, Type II, and Type III
survivorship patterns. Draw a diagram comparing
the three types of survivorship patterns. - How might generation time affect r, the
population growth rate? - Design a study that would allow you to determine
whether a particular species had a more
opportunistic or equilibrial life history. - Think of a habitat near your school or home.
List 10 species that this habitat contains. Do these
species have equilibrial or opportunistic life histories?
Design a study that would allow you to find out. - Compare two species of plants, the dandelion
(Taraxacum officinale) and the red maple (Acer
rubrum) and two species of mammal, such a mouse
and a black bear (Ursus americanus), with respect to
whether they are more r- or K- selected. Specifically,
compare them with respect to their fecundity,
the shape of their survivorship curves, life span,
population density, and whether the environment
each species typically inhabits is more or less stable
or unpredictable.
C
I
I
K/U
K/U
MC
K/U
K/U
C
K/U
K/U
K/U
K/U