THINKING LAB
Metabolic Rate and
the Structure of Molecules
Background
The molecule used by plants, such as corn and potatoes,
to store energy is called starch. Starch is a large polymer
composed of about 1000 glucose molecules. The starch is
formed through condensation reactions, which link together
individual glucose molecules. The starch molecule may
contain side-branches, as shown in the illustration. Before
a plant can use starch in aerobic cellular respiration, the
starch must be broken down into individual glucose
molecules. Recall that glucose is the molecule that first
enters glycolysis. The rate at which glucose is available to
be used in respiration can affect how quickly the cell will
carry out metabolic processes. In other words, the availability
of glucose can determine a plant’s metabolic rate.
You Try It
1.What reaction is needed to break down a starch
molecule into individual glucose molecules?
A starch molecule
2.If sweet corn contains mainly glucose molecules and
starchy corn contains mainly starch molecules, in which
type of corn would you expect cells to have a slower
metabolic rate? Explain briefly.
3.Discuss how the structure of molecules might affect
metabolic rate.
4.How could you determine if corn is sweet or starchy
without tasting it?
5.What is the energy storage molecule in animal cells?
O O O
O O O
O
O
O
O
O
O
O
O
O
O O
O
O
CH 2
92 MHR • Unit 1 Metabolic Processes
Stage 3: Re-formation of RuBP
Recall from Figure 3.27 and Figure 3.28 that RuBP,
ribulose bisphosphate, is required in the carbon
fixation stage of the Calvin cycle. RuBP is used to
produce PGA, needed for the reduction stage of the
cycle. Because PGAL is needed to reform RuBP, the
majority of PGAL molecules, do not contribute to
glucose production. The Calvin cycle reactions
must occur twice to create one molecule of glucose.
This is because for every three times that the
Calvin cycle reactions occur, five PGAL are used to
re-form three RuBP, ribulose bisphosphate, as
shown in Figure 3.32. Notice from Figure 3.27 that
5 three-carbon PGALs contain the same number of
carbon atoms as 3 five-carbon RuBPs.
Figure 3.32Re-formation of RuBP. As five molecules of
PGAL become three molecules of RuBP, three molecules of
ATP become three molecules of ADP+Pi.
To summarize the synthesis reactions of the
Calvin cycle:
Glucose: The Ultimate Food Source
After glucose is produced in the synthesisreactions,
plant cells can use glucose for glycolysis, followed
by aerobic respiration in the mitochondria. The
products and intermediary molecules of aerobic
respiration provide the carbon-based molecules
necessary to build amino acids, as well as the
precursors to nucleic acids and lipids. However,
there are many other ways that plants use glucose,
for example,
the conversion of glucose to starch,
the formation of cellulose from glucose, and
the conversion of glucose to sucrose.
Stage 1: Carbon fixation, which takes carbon
atoms from atmospheric carbon dioxide
molecules and incorporates these atoms
into organic molecules.
Stage 2: Reduction, which involves the
formation of PGAP and its reduction to PGAL.
Stage 3: Re-formation of RuBP, which uses
most of the PGAL molecules formed in the
reduction stage to produce RuBP. This is then
used to form more PGA in the Calvin cycle.
5 PGAL
3 ATP 3 ADP 3
3 RuBP
+ Pi