84 Invasive Stink Bugs and Related Species (Pentatomoidea)
but this scutellar form is seen in many other groups as well. The ostiolar rugae tend to be elongate
and apically acute, curving cephalad. In Eurysaspis, the base of the abdominal venter is armed with
a robust, truncate projection that meets with an elevated metasternum that, in turn, meets with an
elevated mesosternum; the mesosternum becomes narrower and flattened, and protrudes as a rounded
lobe onto the prosternum, between the front coxae (much like that seen in the Rhynchocorini). In
Flaminia, the abdominal venter is not as elevated as in Eurysaspis, but it is still truncately produced
forward and comes into contact with the slightly elevated, but distinctly sulcate, metasternum. The
mesosternum is somewhat flat posteriorly, becomes slightly carinate anteriorly, but is not produced
forward onto the prosternum. In Platacantha, the base of the abdominal venter is armed with a robust,
ventrally flattened spine that extends forward beyond the middle coxae and covers both the meta- and
mesosterna. This is rather similar to the sternal structure exhibited in the Piezodorini, so Platacantha
may need to be transferred to that tribe. In fact, Linnavuori (1982) indicated that Platacantha was
distinctively different from Eurysaspis and should not be placed in the same group with that genus. He
argued that there were some similarities with the genus Acrosternum Fieber (Figure 2.22I) (currently
in the Nezarini), based on the sternal structures, or to members of the Antestiini, based on genitalic
structures.
2.2.10.7.22 Eysarcorini Mulsant and Rey, 1866
Mulsant and Rey (1866) proposed Eysarcoriens as a new family with two branches (= tribes): Eysarcoraires
and Rubiconiaires. Their Rubiconiaires originally contained two genera, Rubiconia and Staria, both
of which are now considered to be members of the Carpocorini (see discussion in Section 2.2.10.7.12).
They originally included three genera in their Eysarcoraires: Dalleria Mulsant and Rey, Eysarcoris
Hahn, and Onylia Mulsant and Rey. Dalleria is currently treated as a subgenus of Stagonomus Gorski,
and Onylia is a junior synonym of the same genus. Two other tribal names have been proposed that are
now considered to be synonyms of the Eysarcorini. Putshkov (1961) proposed the tribe Stolliini, based
on the genus Stollia Ellenrieder, which is considered by most workers to be a synonym of Eysarcoris.
Fuente (1974) proposed the tribal name Stagonomini as a replacement name for Eysarcorini (at one time
there was a dispute concerning the correct type species for Eysarcoris, which could have endangered
the availability of the Eysarcorini). Finally, of minor importance, but worthy of mention is the fact that
the genus Eysarcoris has been unjustifiably emended several times, and, so, various derivations (e.g.,
Eusarcorini, Eysarcocorini) frequent the literature.
This is a difficult tribe to characterize. Its members are usually small to medium in size, and tend to
be somewhat robust in shape (Figure 2.30A). The abdominal venter is unarmed, and the scutellum tends
to be enlarged and spatulate (several exceptions include but are not limited to Aspavia Stål [Figure
2.21G], Carbula Stål [Figure 2.21H], and Durmia Stål). They are usually brown to dark brown in color.
It is a relatively large tribe containing 19 genera and 230 species (Table 2.3). Linnavuori (1982) stated
that the lateral margins of the pronotum are usually callose, whitish, and sharply delimited, except for
the genus Adria (but see further discussion of Adria below). The ostiolar rugae are usually short and
auriculate (Figure 2.8E); the associated evaporative areas are also fairly small. The mesosternum is
medially carinate. In one species of Stagonomus, the female abdomen and genital plates are produced
caudad, becoming acutely pointed posteriorly (Figure 2.21I), much as is seen in the Lestonocorini. The
male parameres are usually strongly F-shaped. The spermathecal bulb is simple, ball-shaped, and lacks
diverticula.
There is little to separate this tribe from the Carpocorini, except for those few genera in which the
scutellum is enlarged. As mentioned above, several included genera do not have the scutellum expanded,
and there are some carpocorine genera (e.g., Coenus) that have an enlarged, spatulate scutellum. The
genus Adria typically has been treated as a member of the tribe Aeliini, but we have tentatively included
it in the Eysarcorini. It may not belong in either tribe. Its species have the mesosternum sulcate, thus
allying it (along with the carpocorine genera Gulielmus and Bonacialus) more closely to the Aeptini,
Diploxyini, Myrocheini, or Sephelini. In fact, Atkinson (1888) treated Adria as a member of the
Diploxyini. Additionally, it is likely that the genus Corisseura Cachan should be transferred to another
tribe. Its members have the ostiolar rugae quite long and acuminate apically. This might place it in the
Cappaeini or Hoplistoderini.