Invasive Stink Bugs and Related Species (Pentatomoidea)

Higher Systematics of the Pentatomoidea 63

14(13) Green in color when alive, and keeping the green color after death (note that some species

have seasonal or semi-permanent yellowish or brownish forms, but the typical form is

green); worldwide (Figures 2.12E; 2.22I-L; 2.30K, L) ........................................Nezarini

14’ Coloration variable, but if predominately green, this green fades to yellow after death .... 15

15(14) Usually small to medium in size; usually tans, browns and/or blacks, sometimes with pale

marks; head relatively broad, usually semi-circular; abdominal spine may be relatively long,

but usually terete, slender; ostiolar rugae elongate, reaching near to lateral margin of meta-

pleuron, apex acute, attached to metapleuron; worldwide (except Europe) (Figures 2.22C-E;

2.30H) ....................................................................................................................Menidini

15’ Usually medium to large in size; color variable, but if tan, brown, or black, then head is not so

broad and/or semicircular; abdominal spine usually large, robust; ostiolar rugae variable .... 16

A

B

C D E

F G H

FIGURE 2.7 A, Mecidea major, abdomen, lateral view; B, Mecidea major, detail of abdominal segments II, III and IV,
lateral view; C, Thyanta acuminata, thorax, ventrolateral view; D, Piezodorus guildiniii, thorax, ventrolateral view; E,
Rhynchocoris humeralis, thorax, ventrolateral view; F, Aelia acuminata, head and thorax, ventral view; G, Aelia acumi-
nata, thorax, ventrolateral view; H, Evoplitus humeralis, thorax, ventrolateral view.