Higher Systematics of the Pentatomoidea 121
2.2.10.9.5 Tarisa Group
The Tar isa group contains three genera and 22 species (Table 2.3) distributed in the Palearctic and
Afrotropical regions.
This group is defined by a specific set or group of characters. They usually have relatively large anten-
niferous tubercles that are partially visible in dorsal view. The anterior part of the pronotum bears tuber-
cles laterally and medially; the scutellum is conspicuously wide basally, completely covering the clavus
and the membrane of the hemelytra; and only part of the corium is exposed. In the male pygophore, the
large part of the ventral wall is either flattened or slightly concave; the infoldings of both the dorsal and
lateral rims bear dense, short setae (Figure 2.10E).
The spermatheca of Dybowskyia reticulata (Dallas) has been described (Kim and Lee 1994). In this
species, the spermathecal bulb is broadly oval with one long process; the distal flange of the pump
region is slightly sclerotized, the proximal flange is not sclerotized, and the region between the flanges
is slightly sclerotized and swollen. The spermathecal duct is slender, proceeding through a sclerotized
dilation; a long proximal part of the duct is free.
All species live on different host plants, Dybowskyia Jakovlev on Apiaceae, and Tar isa on Asteraceae.
Some species of this group are the only known podopines to feed on Chenopodiaceae (recently placed
as a subfamily within the Amaranthaceae) (Schaefer 1981b, 1983; Jitka Vilímová, unpublished data).
2.2.10.9.6 Brachycerocoris Group
The Brachycerocoris group contains three genera and 13 species (Table 2.3) distributed in Palearctic,
Afrotropical, and Oriental regions (Figure 2.18I).
This group of genera is defined by a specific set of characters. In this group, each buccula has a trian-
gular process, either anteriorly or medially (Figure 2.10D). The pronotum has a longitudinal keel either
anteriorly or along its entire length; the scutellum either has distinct tubercles, one basally and one on the
middle in the longitudinal axis, or its basal plate continues as a longitudinal keel. The entire lateral rim
of the pygophore is widened (Jitka Vilímová, unpublished data).
The spermatheca of Bolbocoris variolosus (Germar) has been illustrated (Linnavuori 1982). The sper-
mathecal bulb is globular; the two flanges of the pumping region are of similar size and shape, and
the region between the flanges is relatively long, longer than the spermathecal duct before it enters the
sclerotized rod.
The members of the genus Bolbocoris Amyot and Serville are known only from Poaceae, whereas
species of the genus Brachycerocoris Costa occur on a wide spectrum of plants including Fabaceae,
Rhamnaceae, and Verbenaceae (Schaefer 1981b, 1983; Linnavuori 1982). Göllner-Scheiding (1992) pro-
vided a revision of the genus Bolbocoris, and Schaefer et al. (1996) revised the genus Brachycerocoris.
2.2.10.9.7 Genera incertae sedis
The six genera listed below are classified as incertae sedis. They do not show any characteristics (char-
acter or group of characters) that would enable us to include them in any of the monophyletic groups
mentioned above. The distribution of most of the genera are summarized by Schouteden (1905b).
Cryptogamocoris Carapezza, 1997
This genus has a single species (C. cornutus Carapezza) known only from Tunisia where it lives on
Salsola vermiculata L. (Amaranthaceae). In general appearance, this species is similar to Tar isa
(Carapezza 1997), but there also appears to be some similarities with members of the Graphosoma
group. Because the specific and important characters of the pygophore that distinguish these two groups
have not been described, we cannot confidentally resolve its proper placement.
Cyptocoris Burmeister, 1835
This genus, distributed in the Afrotropical region, has been included in the Graphosomatini by sev-
eral authors (Schouteden 1905b, Gillon 1972, Linnavuori 1982). However, its relationship has not been
resolved. In fact, several characters are not congruent with its placement in the Podopinae. Its compound
eyes are not prominent, but, instead, they lie within the outline of the head. The head is strongly convex
dorsally, almost semiglobular. The frena are not developed, and the pygophoral area is extremely narrow.