A CENTURY OF PROGRESS 147
fossils he found on his explorations,
which he now saw “through Lyell’s
eyes.” However, while he was in
South America, volume two of
Principles of Geology arrived. In
it, Lyell rejected ideas of gradual
evolution of plants and animals,
including Lamarck’s theories.
Instead, he invoked the concept
of “centres of Creation” to explain
species’ diversity and distribution.
Although Darwin admired Lyell as
a geologist, he had to discount this
latest concept as the evidence for
evolution mounted.
Another piece of the jigsaw
was revealed in 1838 when Darwin
read An Essay on the Principle
of Population by the English
demographer Thomas Malthus,
which had been published 40 years
earlier. Malthus described how
human populations can increase
in an exponential way, with the
potential to double after one
generation of 25 years, then double
again in the next generation, and
so on. However, food supplies
could not expand in the same way,
and the result was a struggle for
existence. Malthus’s ideas were
one of the main inspirations for
Darwin’s theory of evolution.
The quiet years
Even before the Beagle had
returned to England, the interest
generated by the specimens
Darwin had sent back had made
him a celebrity. After his return,
his scientific and popular accounts
of the voyage increased his fame.
However, his health deteriorated
and gradually he withdrew from
the public eye.
In 1842, Darwin moved to the
peace and quiet of Down House in
Kent, where he continued to amass
evidence to support his theory of
evolution. Scientists around the
world sent him specimens and
data. He studied the domestication
of animals and plants, and the
role of selective breeding, or artificial
selection, especially in pigeons. In
1855, he started breeding varieties
of Columbia livia, or rock doves, and
they would feature prominently in
the first two chapters in On the
Origin of Species.
Through his work on pigeons,
Darwin began to understand
the extent and relevance of
variation among individuals.
He rejected the accepted wisdom
that environmental factors were
responsible for such differences,
insisting that reproduction was
the cause, with variation somehow
inherited from parents. He added
this to the ideas of Malthus and
applied them to the natural world. ❯❯
The finches of the Galápagos have evolved
differently shaped beaks adapted to specific diets.
Geospiza
magnirostris
Camarhynchus
parvulus
Geospiza fortis
Certhidea
olivacea
Medium-sized
ground finch has
a smaller beak for
crushing smaller,
softer seeds.
Warbler finch
has a thin beak for
probing for small
insects and
spearing them.
Large ground
finch has a large,
strong beak for
crushing large,
woody seeds.
Small tree finch
has a short, sharp
beak for grasping
insects.